biology major phd

Ph.D. Program

The training for a Ph.D. in Biology is focused on helping students achieve their goals of being a successful research scientist and teacher, at the highest level. Students work closely with an established advisor and meet regularly with a committee of faculty members to facilitate their progress. The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. 

There are two tracks within the Biology Ph.D. program:

• Cell, Molecular and Organismal Biology
• Ecology and Evolution

(Previously a part of the Department of Biology Hopkins Marine Station is now a part of the Oceans Department within  Stanford Doerr School of Sustainability )

All  tracks are focused on excellence in research and teaching in their respective areas; where there are differences between the tracks, they are indicated in the links below. 

Requirements & Forms

Dissertation defense, cellular and molecular biology training program, stanford biology ph.d. preview program, career development resources.

Biology, PHD

On this page:, at a glance: program details.

• Location: Tempe campus
• Second Language Requirement: No

Program Description

Degree Awarded: PHD Biology

The PhD program in biology offers individualized courses of study tailored to students' interests that include laboratory, field and theoretical work. Flexibility in the program is achieved by requiring only one core class, which is a choice between two topics that cover the breadth of the research directions in this program.

Electives are also as flexible as possible and include areas outside of biology. The major goal is to provide students with the opportunity to create an adaptable and general degree program that allows both traditional and interdisciplinary approaches in any area of biology.

Program Faculty   PhD Students

• BIO 792 Research
• BIO 514 Statistical Models for Biology
• BIO 522 Populations: Evolutionary Ecology
• BIO 507 Advanced Evolutionary Medicine
• BIO 521 Landscape Ecology
• BIO 530 Scientific Teaching (required in first semester for students that will TA at any point in degree)
• BIO 533 K-12 STEM Education & Outreach
• BIO 539 Computing for Research
• BIO 541 SOLS Seminar Series
• BIO 542 SOLS Current Topics in the Life Sciences
• BIO 591 Society and Natural Resource Management
• BIO 591 Social-Ecological Systems & Adaptation
• BIO 591 Drylands in a Changing Earth
• BIO 591 Population Genetic Reading Group
• BIO 591 Communication for Scientists
• BIO 591 Genetics and Genomics of Behavior
• BIO 620 Research Prospectus Writing
• EVO 501 Current Topics in Evolutionary Biology
• EVO 601 Principles of Evolution
• EVO 610 Research Areas of Evolution

Application and admission information

How to apply.

Applications open September 1 for admission in Fall of the following year. The application deadline is December 1 . We accept applications for Fall semesters only. We cannot guarantee that applications received after the December 1 deadline will be considered for admission.

All applicants must apply by filling out ASU's Graduate Admissions application. All application materials must be submitted through the application or to Graduate Admissions directly. Please do not mail or email any documents to the School of Life Sciences. 

Required materials and information include the following:

• 1-2 page personal statement
• An up to date CV or resume
• The names of relevant SOLS faculty you have been in touch with who you might be interested in being supervised by
• Unofficial transcripts and English proficiency test scores (if applicable)
• The names and emails of at least 3 recommenders to write you letters of recommendation

Application review process and timeline

Following the December 1 deadline, faculty will begin reviewing applications. Applicants should monitor their My ASU priority tasks to ensure there are no missing materials in their application.

Faculty will decide which applicants they would like to invite to our Graduate Recruitment Weekends (GRWs), typically held in February. Applicants will hear from the School of Life Sciences in January if they are invited to participate in the GRWs.

Admission decisions will begin after the GRWs, and applicants typically receive final decisions by April 1.

Requirements

Minimum requirements for admission include the following:

• Cumulative GPA of at least 3.0 on a 4.0 scale
• There are other ways to demonstrate English proficiency beyond the tests, so please refer to ASU's English proficiency webpage to review how you might satisfy requirements

Desired qualifications typically seen in competitive candidates:

• Research experience and a letter of recommendation from a faculty research supervisor
• English proficiency scores that meet these teaching assistant language proficiency requirements

Please note that the GRE is not required.

Students offered admission to a PhD program in the School of Life Sciences will typically receive a funding offer as well. While individual funding offers may differ to some degree, they typically include teaching assistant and/or research assistant positions each semester (summer optional) for 5 years. These positions provide financial coverage through the following:

• A standard salary stipend paid biweekly
• Tuition remission covering enrollment in 6-18 credit hours for fall and spring semesters and 1-14 credit hours for summer semesters
• Health insurance coverage

To discover more, check out the ASU Graduate College's funding opportunities !

Degree Requirements

84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation

Required Core (3 or 4 credit hours) BIO 514 Statistical Models for Biology (4) or BIO 620 Research Prospectus Writing (3)

Electives (67 or 68 credit hours)

Other Requirements (1 credit hour) BIO 610 Introduction to Responsible Conduct of Research in Life Sciences (1)

Culminating Experience (12 credit hours) BIO 799 Dissertation (12)

Additional Curriculum Information The program is tailored to the needs of the individual student. The plan of study is developed by the student and a supervisory committee consisting of a major professor and three additional faculty members. The electives credit hours are comprised of coursework showing breadth in biology subdisciplines, seminars and research.

An oral comprehensive examination is required in order to advance to candidacy, and a formal defense of the final written dissertation is required prior to graduation.

Admission Requirements

Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.

Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in biology or a related discipline from a regionally accredited institution.

Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program, or a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in an applicable master's degree program.

Applicants must submit the following:

• graduate admission application and application fee
• official transcripts
• academic record form
• personal statement
• curriculum vitae or resume
• three letters of recommendation
• proof of English proficiency

Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.

Research experience is desired for entry into this program.

Next Steps to attend ASU

Learn about our programs, apply to a program, visit our campus, learning outcomes.

• Able to communicate the findings and originality of their research, both orally and in writing.
• Able to perform research that they have designed to address important and challenging questions in the field of biology.
• Able to review scholarly literature as they develop specialized expertise in their selected research area.

Career Opportunities

Those who have earned a doctorate in biology are prepared for academic careers at every level, from community colleges to research universities. Their skills and knowledge are also valuable for government careers with federal and state agencies responsible for wildlife management and conservation, and for careers in industry and nongovernmental organizations.

Career examples include:

• food, agriculture and health care scientists in academic, private and industrial labs
• principal investigators in government labs and nonprofit organizations
• professors or instructors in universities and colleges
• science teachers in elementary and high schools
• wildlife, animal and conservation scientists

Program Contact Information

If you have questions related to admission, please click here to request information and an admission specialist will reach out to you directly. For questions regarding faculty or courses, please use the contact information below.

• [email protected]
• 480/965-1768

PhD Program in Biology

Our graduate program provides students of diverse backgrounds with the opportunity to intensely engage in research in the biological sciences and contribute to the broader scientific community.

Average time to degree: 5.3 years

Percentage of graduates in post-doctoral or permanent positions: 94%

Average publications after completion of program: 3.8 per person

PhD Program

The Department of Biology introduces graduate students to diverse fields of biological science, and provides them with expert guidance to excel in research. The department is invested in training students to become excellent scientists, researchers, science communicators, and instructors. We are a diverse and global community, committed to expanding scientific career opportunities to all. Some of our graduates become academics, whereas others find careers in government, private industry, public policy, or elsewhere (see where recent graduates are now). The ability to communicate ideas and research results clearly and convincingly is key to success in any career.

Experience Biology at Georgetown University

Professors and current students share their perspective of Georgetown’s cutting edge biology graduate program.

Potential applicants are urged to identify and contact potential research mentors directly before applying. Please refer to the research page and the list of faculty interested in accepting students . Not all laboratories will have open positions available for a new graduate student in a given year.

Financial Assistance for PhD Students

    students who are accepted into the phd program in biology are guaranteed a minimum of five years of full funding. this funding includes:.

Stipend The stipend for PhD students in Biology is set by the Collective Bargaining Agreement between Georgetown University and the Georgetown Alliance of Graduate Employees ( GAGE ). The current stipend rate is $36,934 for the year. The stipend is paid on a 12-month contract, typically in 26 biweekly pay checks.

Tuition Waiver The Graduate school of Arts & Sciences (GSAS) provides full tuition waivers for all PhD students for the duration of the PhD program.

Health Insurance Health insurance is required. The GSAS provides free student health insurance to all students. The insurance covers doctor’s visits, hospital stays, and prescription drugs. Students may waive this benefit if they prefer a different plan through parents or a spouse.

Dental Insurance The GSAS provides free dental insurance to all students. The insurance covers routine dental care, such as cleanings and exams.

Yates Athletic Facility All students are eligible to use the Yates athletic facility for free. The facility has a gym, swimming pool, and fitness classes.

Parental Leave Graduate student workers are guaranteed six weeks of paid parental leave. This leave can be used to bond with a new child or to care for a sick child.

Medical Leave Graduate student workers are guaranteed six weeks of paid medical leave. This leave can be used to recover from a medical condition or to care for a sick family member.

To learn more about financial assistance for PhD students, please visit the Graduate Funding page. You can read more about what it means to do a PhD in a union-protected graduate program  here

Student Quotes

“Until you start your journey in grad school you might not know what you are missing out on. Other than the course work, grad school has been instrumental in  inculcating important life values and skills such as confidence, time management, survival, and holding personal and professional relationships in higher esteem. ”

“Studying at Georgetown has enabled me to be a lot more confident in my ability to ask questions and chart my own path in scientific research. My advisor and mentors here at Georgetown have been top quality in terms of both academic and personal support”

“Georgetown is a place with an environment conducive to learning and professional growth for prospective scientists working towards their graduate degrees and beyond. In this program, I found mentorship and training that foster the critical thinking required to answer the questions I am interested in. I found the multidisciplinary nature of the graduate student body to be engaging and supportive from the beginning of my journey.”

“The breadth of research that goes on in the Bio department really facilitates a broad understanding of biology from the life of single cells to the life of much larger organisms. It really is something unique that you don’t get at other more specific departments and has really helped me think outside of the box with my own work.”

Graduate Research Opportunities

Learn about the research programs that the department of Biology offers and find out which programs are recruiting graduate students.

Graduate Teaching Opportunities

Find out about the many opportunities that are offered to graduate students to develop teaching and mentoring skills.

Graduate Funding Opportunities

Learn about stipend funding and the additional funds from the GU Graduate School offered to help support graduate dissertation research or travel to meetings.

PhD in Biology

The PhD program in Biology is a research-intensive program that also has a strong focus on teaching, designed to produce top scientists and educators with a broad base of knowledge to tackle the most important biological problems of today. This is accomplished through research training, graduate-level courses, seminars, and teaching experience. Entering students are assigned an advisory committee of three faculty members who work with the student to plan a suitable program based on the student's experience and interests.

PhD students in Biology receive full tuition support for six years, a competitive stipend, health coverage, and receive extensive training in pedagogy, outreach, and communication that prepares them for careers in academia, biotechnology, education, and policy. Students are required to serve as teaching assistants for at least two semesters. Most students complete the requirement for teaching experience in the first year by assisting in the introductory undergraduate biology courses for two semesters.

During the first year, students become familiar with research opportunities in the department by doing short research rotations with various faculty members. This experience in expected to lead to the selection of a research focus for the PhD thesis, and it also sets the stage for cross-disciplinary approaches to the thesis topic that is pursued.

First- and second-year students usually take some graduate courses that are selected in consultation with the advisory committee, in addition to one required course: either Biology 243: Topics in Molecular and Cell Biology; Biology 244: Topics in Evolutionary Ecology; or Biology 246: Topics in Physiology and Animal Behavior.

The department faculty are educators as well as researchers, and the graduate students benefit from this expertise. Graduate students serving as teaching assistants have the opportunity to take a course on teaching and pedagogy to improve their skills, or to become involved with upper level courses in their area of expertise. In addition, we offer a one-month summer program for PhD students interested in contributing to course design and honing their teaching skills via GIFT, the Graduate Institute For Teaching .

Research Concentrations

Browse the six concentration areas below for recommended programs of study and relevant courses:

• Biology Education Research
• Ecology, Behavior and Evolution
• Global Change Biology
• Genetics and Molecular Biology
• Developmental Biology
• Physiology, Neurobiology and Animal Behavior

Graduate credit for a course requires a grade of B- (B minus) or better. A list of biology department undergraduate and graduate courses can be found by selecting the Courses button from the top bar.

Throughout the year, graduate students benefit from a variety of seminars on current research that are presented by faculty, fellow graduate students, and invited speakers.

Biology Graduate Programs in America

1-25 of 396 results

MIT School of Science

Cambridge, MA •

Massachusetts Institute of Technology •

Graduate School

Massachusetts Institute of Technology ,

Graduate School ,

CAMBRIDGE, MA ,

Stanford University Department of Humanities and Sciences

Stanford, CA •

Stanford University •

Stanford University ,

STANFORD, CA ,

Stanford University School of Medicine

• • Rating 5 out of 5   3 reviews

Doctoral Student: I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. ... Read 3 reviews

3 Niche users give it an average review of 5 stars.

Featured Review: Doctoral Student says I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. .

Read 3 reviews.

University of North Texas

Graduate School •

• • Rating 4.61 out of 5   129

College of Arts and Sciences - American University

American University •

WASHINGTON, DC

Lehigh University

BETHLEHEM, PA

• • Rating 4.45 out of 5   20

Harvard Graduate School of Arts and Sciences

Harvard University •

• • Rating 4.56 out of 5   9 reviews

Other: I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student' in the Harvard Graduate School of Arts and Sciences. Through this School, I will be focusing my time at the John A. Paulson school of Engineering & Applied Sciences. Looking forward to wrapping up my final year on campus! ... Read 9 reviews

Harvard University ,

9 Niche users give it an average review of 4.6 stars.

Featured Review: Other says I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student'... .

Read 9 reviews.

Harvard Medical School

Boston, MA •

• • Rating 4.94 out of 5   16 reviews

Other: Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try my best of the best….. ... Read 16 reviews

BOSTON, MA ,

16 Niche users give it an average review of 4.9 stars.

Featured Review: Other says Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try... .

Read 16 reviews.

Duke University School of Medicine

Durham, NC •

Duke University •

• • Rating 4.76 out of 5   17 reviews

Doctoral Student: Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. ... Read 17 reviews

Duke University ,

DURHAM, NC ,

17 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. .

Read 17 reviews.

• Find college scholarships

Brown University Graduate School

Providence, RI •

Brown University •

Brown University ,

PROVIDENCE, RI ,

School of Arts & Sciences - University of Pennsylvania

Philadelphia, PA •

University of Pennsylvania •

University of Pennsylvania ,

PHILADELPHIA, PA ,

Weinberg College of Arts and Sciences

Evanston, IL •

Northwestern University •

Northwestern University ,

EVANSTON, IL ,

Vanderbilt University School of Medicine

Nashville, TN •

Vanderbilt University •

• • Rating 5 out of 5   2 reviews

Graduate Student: Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. ... Read 2 reviews

Vanderbilt University ,

NASHVILLE, TN ,

2 Niche users give it an average review of 5 stars.

Featured Review: Graduate Student says Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. .

Read 2 reviews.

Washington University in St. Louis - Arts & Sciences

St. Louis, MO •

Washington University in St. Louis •

Washington University in St. Louis ,

ST. LOUIS, MO ,

Guarini School of Graduate and Advanced Studies

Hanover, NH •

Dartmouth College •

• • Rating 5 out of 5   1 review

Alum: I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary learning. Guarini is well integrated within the College, which provides a unique world-class learning experience. Guarini went beyond my expectations and made this experience the best I had and will ever have in my life. The only thing I would like changed is having a GRAD diploma in Latin instead of English to keep up with College tradition. ... Read 1 review

Dartmouth College ,

HANOVER, NH ,

1 Niche users give it an average review of 5 stars.

Featured Review: Alum says I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary... .

Read 1 reviews.

• Sponsored Find Student Loan Options
• Biostatistics Graduate Programs
• Computational Biology Graduate Programs

Division of Biology and Biological Engineering - California Institute of Technology

Pasadena, CA •

California Institute of Technology •

California Institute of Technology ,

PASADENA, CA ,

College of Science - University of Notre Dame

Notre Dame, IN •

University of Notre Dame •

University of Notre Dame ,

NOTRE DAME, IN ,

Biological Sciences Division - University of Chicago

Chicago, IL •

University of Chicago •

• • Rating 4 out of 5   1 review

Master's Student: So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. ... Read 1 review

University of Chicago ,

CHICAGO, IL ,

1 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. .

Pritzker School of Medicine

Dornsife college of letters, arts and sciences.

Los Angeles, CA •

University of Southern California •

University of Southern California ,

LOS ANGELES, CA ,

Cornell University College of Arts & Sciences

Ithaca, NY •

Cornell University •

Cornell University ,

ITHACA, NY ,

Georgetown University School of Medicine

Nw Washington, DC •

Georgetown University •

• • Rating 4.75 out of 5   4 reviews

Doctoral Student: Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1 and Step 2 preparation. ... Read 4 reviews

Georgetown University ,

NW WASHINGTON, DC ,

4 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1... .

Read 4 reviews.

School of Arts and Sciences - Tufts University

Medford, MA •

Tufts University •

• • Rating 4 out of 5   4 reviews

Master's Student: My experience was mostly great! I felt supported by faculty and staff and became involved in student activities. It was just unfortunate that my on-campus experience was cut short due to the pandemic so I was not able to fully take advantage of all the program and graduate school had to offer. ... Read 4 reviews

Tufts University ,

MEDFORD, MA ,

4 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says My experience was mostly great! I felt supported by faculty and staff and became involved in student activities. It was just unfortunate that my on-campus experience was cut short due to the pandemic... .

UCLA College of Letters and Science

University of California - Los Angeles •

• • Rating 3 out of 5   1 review

University of California - Los Angeles ,

1 Niche users give it an average review of 3 stars.

Krieger School of Arts & Sciences

Baltimore, MD •

Johns Hopkins University •

• • Rating 4.53 out of 5   19 reviews

Master's Student: I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student Advisor, Alexis has been extremely helpful in initiating my enrollment process and answering all of my questions in a timely matter. My intended Advanced Academic Program is the accelerated (2 semester), dual-modality, 40-credit M.S. in Biotechnology, Biodefense concentration. All of the anticipated course subjects are diverse and there's even a customizable core lab course on campus (at least until Summer 2024). I can't wait and I wish you all the best in your search for academic programs or professional certifications. ... Read 19 reviews

Johns Hopkins University ,

BALTIMORE, MD ,

19 Niche users give it an average review of 4.5 stars.

Featured Review: Master's Student says I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student... .

Read 19 reviews.

Mellon College of Science

Pittsburgh, PA •

Carnegie Mellon University •

Blue checkmark.

Carnegie Mellon University ,

PITTSBURGH, PA ,

The Graduate School of Arts & Sciences - University of Virginia

Charlottesville, VA •

University of Virginia •

Alum: Very good in some areas, excellent in other areas, many academic choices available in all areas of study ... Read 1 review

University of Virginia ,

CHARLOTTESVILLE, VA ,

Featured Review: Alum says Very good in some areas, excellent in other areas, many academic choices available in all areas of study .

College of Sciences - Georgia Institute of Technology

Atlanta, GA •

Georgia Institute of Technology •

Other: Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer to newcomers is to prioritize building friendships early on. One regret I have as I reflect on my year is not connecting with more people. I initially thought I should always work independently, but that was a misstep in my academic journey. Remember, it's essential to strike a balance between learning and having fun. ... Read 1 review

Georgia Institute of Technology ,

ATLANTA, GA ,

Featured Review: Other says Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer... .

Walla Walla University

COLLEGE PLACE, WA

• • Rating 4.36 out of 5   14

Florida International University

• • Rating 4.64 out of 5   296

Mississippi State University College of Arts and Sciences

Mississippi State University •

MISSISSIPPI STATE, MS

Showing results 1 through 25 of 396

Biological and Biomedical Sciences

Share this page.

Biological and Biomedical Sciences (BBS)  is an area of study within the Division of Medical Sciences, an administrative unit based at Harvard Medical School that coordinates biomedical PhD activities at the Longwood Medical Area. Students who study in BBS receive a PhD in medical sciences. Prospective students apply through the Harvard Kenneth C. Griffin Graduate School of Arts and Sciences (Harvard Griffin GSAS). In the online application, select  “Division of Medical Sciences” as your program choice and select "Biological and Biomedical Sciences" in the area of study menu.

This program trains you in the biosciences, starting with core training in contemporary genetics; biochemistry; and molecular, cellular, and mechanistic biology. You can customize your curriculum to allow you to align with your research interests.

You will have access to a vast wealth of resources, including more than 700 faculty, top scientists, core facilities and hospitals, cutting-edge research, and an extensive alumni network. You will be in close proximity to top nationwide hospitals and research centers and more than 130 biotech, biopharma, and pharmaceutical companies that have biological and biomedical sciences (BBS) alumni on staff.

Examples of specific student projects include “Development of nucleic acid detection methods for object provenance and viral diagnostics,” “Interrogating genetic diversity in Mycobacterium abscessus with transposon-sequencing,” and “Sexual dimorphism in hair follicle stem cells.”

Graduates of the program have secured faculty positions at such prestigious institutions as New York University, University of Minnesota, Harvard University, and University of Illinois. Others have gone on to careers with leading companies such as Novartis and the Broad Institute. Derrick Rossi, cofounder of Moderna, is a graduate of the BBS program.

Standardized Tests

GRE General: Not Accepted GRE Subject: Not Accepted iBT TOEFL minimum score: 100 IELTS minimum score: 7

See list of Biological and Biomedical Sciences faculty

APPLICATION DEADLINE

Questions about the program.

Secondary Menu

Graduate program.

Duke Biology offers diverse training opportunities for students seeking a Ph.D. degree in biological sciences. Our program is highly flexible, allowing students to design the path that best fits their professional needs. Our students spend the majority of their time engaged in innovative research, and have the opportunity to interact and collaborate with researchers from a wide range of fields. 

The graduate students in the Biology Department (Biograds) are integral to the department and community. Contributions include: running weekly happy hours and prospective student recruitment weekends as well as organizing seminars, journal clubs, teaching-assistant training orientation, and diversity-themed workshops.

Program Requirements

Requirements for the Ph.D. include course work, participation in seminars relevant to your field of research, successful completion of an oral qualifying examination, one year of teaching experience, completion and defense of a research dissertation under faculty guidance.

Affiliated Graduate Programs

Duke Biology is involved in multiple University-wide graduate programs, which link researchers across different departments and schools at Duke – most notably the basic sciences departments in the  Duke University Medical Center  and the  Nicholas School of Earth and Ocean Sciences .

For Prospective Students

Learn more about our degree, our university, and our community. We'll explain the application process, financial support, and more.

• For Current Students

We've compiled resources to assist you while working toward your Ph.D. – from requirements and fellowships, to milestones, useful Duke links, and more.

Learn more about the application process, deadlines and other resources for our candidates.

Review Application Information

• Duke Biology’s Mission Statement
• AJED Annual and Semester Reports
• AJED Meeting Notes
• Biology Cultural Association (BCA)
• Inclusion, Diversity, Equity, and Antiracism Committee (IDEA)
• Learning from Baboons: Dr. Susan Alberts
• Extremophiles and Systems Biology: Dr. Amy Schmid
• How Cells Manage Stress: Dr. Gustavo Silva
• Predator-Prey Interactions in a Changing World: Dr. Jean Philippe Gibert
• Exploring the Extracellular Matrix: Dr. David Sherwood
• Cell Division's Missing Link: Dr. Masayuki Onishi
• Listening in to Birdsong: Dr. Steve Nowicki
• Biogeochemistry as Ecosystem Accounting: Dr. Emily Bernhardt
• Building a Dynamic Nervous System: Dr. Pelin Volkan
• Investigating a Key Plant Hormone: Dr. Lucia Strader
• Imagining Visual Ecology: Dr. Sönke Johnsen
• Outreach Opportunities Across the Triangle
• Job Opportunities
• Location & Contact
• Frequently Asked Questions
• Learning Outcomes
• Major Requirements
• Anatomy, Physiology & Biomechanics
• Animal Behavior
• Biochemistry
• Cell & Molecular Biology
• Evolutionary Biology
• Marine Biology
• Neurobiology
• Pharmacology
• Plant Biology
• Minor Requirements
• Biology IDM
• List of Biology Advisors
• Guide for First-Year Students
• Transfer Credit
• Application & Deadlines
• Supervisor & Faculty Reader
• Thesis Guidelines
• Honors Poster
• Past Student Projects
• Study Away Opportunities
• Finding a Research Mentor
• Project Guidelines
• Getting Registered
• Writing Intensive Study
• Independent Study Abroad
• Summer Opportunities
• Departmental Awards
• Biology Majors Union
• Commencement 2024
• Trinity Ambassadors
• Degree Programs
• Ph.D. Requirements
• How to Apply
• Financial Aid
• Living in Durham
• Where Our Students Go
• Milestones Toward Ph.D.
• Graduate School Fellowships
• Useful Resources
• Concurrent Biology Master of Science
• En Route Biology Masters of Science
• Form Library
• Mentorship Expectations
• On Campus Resources
• Fellowships & Jobs
• Meet Our Postdocs
• Department Research Areas
• Research Facilities
• Duke Postdoctoral Association
• All Courses
• Biological Structure & Function Courses
• Ecology Courses
• Organismal Diversity Courses
• Alternate Elective Courses
• Primary Faculty
• Secondary Faculty
• Graduate Faculty
• Emeritus Faculty
• Graduate Students
• Department Staff
• Faculty Research Labs
• Developmental Biology
• Ecology & Population Biology
• Neuroscience
• Organismal Biology & Behavior
• Systematics
• Research Articles & Papers
• Botany Plot
• Field Station
• Pest Management Protocols
• Research Greenhouses
• Centers/Research Groups
• Biology Writes
• Alumni Profiles
• Assisting Duke Students

• Skip to Content
• Catalog Home
• Institution Home
• Graduate Catalog /
• School of Arts & Sciences /

Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2023 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

Sample Plan of Study

Print options.

Print this page.

The PDF will include all information unique to this page.

A PDF of the entire 2023-24 catalog.

A PDF of the 2023-24 Undergraduate catalog.

A PDF of the 2023-24 Graduate catalog.

Boston University Academics

Boston University

• Campus Life
• Schools & Colleges
• Degree Programs
• Search Academics
• PhD in Biology

The goal of the Biology Department is to train students at the highest level in one of three broad subfields within modern biology: Cell & Molecular Biology, Neurobiology, and Ecology, Behavior, Evolution & Marine Biology. Upon completion of the PhD, students should be prepared for postgraduate training and to eventually assume teaching and/or research positions in academia, industry, government, or nonprofit agencies. The PhD is a research degree and normally necessitates at least five years of academic study, including summer work.

Applicants to the PhD program must have completed a bachelor’s degree in biology or a closely related field. We favor applicants with both strong academic records and a demonstrated aptitude for research.

Learning Outcomes

• Demonstrate academic mastery in one of three areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.
• Attain research expertise, including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.
• Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.
• Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Course Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits. See the GRS Transfer of Credits policy for more details.

• 2 semesters of Progress in Research Seminars (1–2 credits each, select from CAS BI 583, BI 584, BI 579, BI 580, GRS NE 500, NE 501)
• 1 semester grant-writing course (2 credits, usually GRS BI 671 or CAS BI 581)
• 1 semester pedagogy course (1 credit, BI 697)
• 1 quantitative course (3–4 credits, selected from a list of courses)
• Additional courses vary by specific track and student interests

Cell & Molecular Biology

• GRS BI 735 Advanced Cell Biology
• GRS BI 753 Advanced Molecular Biology
• GRS MB 721 Graduate Biochemistry
• Three electives
• Research credits

Neurobiology

• GRS BI 755 Cellular and Systems Neuroscience
• GRS BI 756 Systems and Behavioral Neuroscience or BI 741 Neural Systems: Functional Circuit Analysis
• Four electives

Ecology, Behavior, Evolution & Marine Biology

• Six electives

Language Requirement

There is no foreign language requirement for this degree.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the Cell & Molecular Biology and Ecology, Behavior, Evolution & Marine Biology curricula, this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation and Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge. The course provides guidance and training on pedagogy and other aspects of graduate school.

MS for PhD Students (Two Options)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam) and has completed 32 credits of graduate-level coursework (not including research) may apply to the graduate school for a Master of Science (MS) degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two:  A PhD student who has taken, but has not advanced to candidacy based on, the PhD qualifying examination may still receive a master’s degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for an MS degree and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Related Bulletin Pages

• Certificate in Biogeoscience
• Graduate School of Arts & Sciences Departments
• Graduate School of Arts & Sciences Courses
• Abbreviations and Symbols

Beyond the Bulletin

• Department of Biology
• Graduate School of Arts & Sciences
• Graduate Admissions
• Graduate Financial Aid
• BA/MA Program
• Master’s Degree Requirements
• PhD Degree Requirements
• African American Studies
• American & New England Studies
• Anthropology
• Archaeology
• Bioinformatics
• MS in Biology
• Biostatistics
• Classical Studies
• Cognitive & Neural Systems
• Computer Science
• Creative Writing
• Earth & Environment
• Editorial Studies
• History of Art & Architecture
• Latin American Studies
• Linguistics
• Literary Translation
• Mathematics & Statistics
• Molecular Biology, Cell Biology & Biochemistry
• Neuroscience
• Pardee School of Global Studies
• Playwriting
• Political Science
• Preservation Studies
• Religious Studies
• Romance Studies
• Sociology & Social Work
• Statistical Practice
• African Studies Certificate
• Asian Studies Certificate
• Advanced Biogeoscience Certificate
• Holocaust, Genocide & Human Rights Studies Certificate
• Latin American Studies Certificate
• Linguistics Certificate
• Museum Studies Certificate
• Muslim Studies Certificate
• Teaching Language, Literature & Film Certificate
• Teaching Writing Certificate
• Women’s, Gender & Sexuality Studies Certificate
• Departments
• Research Centers & Institutes

Terms of Use

Note that this information may change at any time. Read the full terms of use .

related websites

Accreditation.

Boston University is accredited by the New England Commission of Higher Education (NECHE).

• © Copyright
• Mobile Version

Biology, PhD

• Program description
• At a glance
• Degree requirements
• Admission requirements
• Tuition information
• Program learning outcomes
• Career opportunities
• Contact information

Science, Scientist, approved for STEM-OPT extension, cell, molecular, plant, sols

You can have the largest amount of flexibility in your course choices in a program that complements other, more specialized life science degree programs. Develop your research skills and acquire high research competency in one or more specialized areas while receiving a broad, solid grounding in biological sciences.

The PhD program in biology offers individualized courses of study tailored to students' interests that include laboratory, field and theoretical work. Flexibility in the program is achieved by requiring only one core class, which is a choice between two topics that cover the breadth of the research directions in this program.

Electives are also as flexible as possible and include areas outside of biology. The major goal is to provide students with the opportunity to create an adaptable and general degree program that allows both traditional and interdisciplinary approaches in any area of biology.

This program may be eligible for an Optional Practical Training extension for up to 36 months. This OPT work authorization term may help international students gain skills and experience in the U.S. Those interested in an OPT extension should review ASU degrees that qualify for the STEM-OPT extension at ASU's International Students and Scholars Center website.

The OPT extension only applies to students on an F-1 visa and does not apply to students completing the degree through ASU Online.

• College/school: The College of Liberal Arts and Sciences
• Location: Tempe

84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation

Required Core (3 or 4 credit hours) BIO 514 Statistical Models for Biology (4) or BIO 620 Research Prospectus Writing (3)

Electives (67 or 68 credit hours)

Other Requirements (1 credit hour) BIO 610 Introduction to Responsible Conduct of Research in Life Sciences (1)

Culminating Experience (12 credit hours) BIO 799 Dissertation (12)

Additional Curriculum Information The program is tailored to the needs of the individual student. The plan of study is developed by the student and a supervisory committee consisting of a major professor and three additional faculty members. The electives credit hours are comprised of coursework showing breadth in biology subdisciplines, seminars and research.

An oral comprehensive examination is required in order to advance to candidacy, and a formal defense of the final written dissertation is required prior to graduation.

Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.

Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in biology or a related discipline from a regionally accredited institution.

Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program, or a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in an applicable master's degree program.

Applicants must submit the following:

• graduate admission application and application fee
• official transcripts
• academic record form
• personal statement
• curriculum vitae or resume
• three letters of recommendation
• proof of English proficiency

Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.

Research experience is desired for entry into this program.

Program learning outcomes identify what a student will learn or be able to do upon completion of their program. This program has the following program outcomes:

• Able to review scholarly literature as they develop specialized expertise in their selected research area.
• Able to perform research that they have designed to address important and challenging questions in the field of biology.
• Able to communicate the findings and originality of their research, both orally and in writing.

Those who have earned a doctorate in biology are prepared for academic careers at every level, from community colleges to research universities. Their skills and knowledge are also valuable for government careers with federal and state agencies responsible for wildlife management and conservation, and for careers in industry and nongovernmental organizations.

Career examples include:

• food, agriculture and health care scientists in academic, private and industrial labs
• principal investigators in government labs and nonprofit organizations
• professors or instructors in universities and colleges
• science teachers in elementary and high schools
• wildlife, animal and conservation scientists

School of Life Sciences | LSA 181 [email protected] 480-965-1768

Department of Biology

• Majors & Minors
• Graduate Programs
• Faculty & Research
• Financial Aid
• Ph.D. Program

The Department of Biology offers a Doctor of Philosophy degree in Quantitative Biology with research emphasis in a number of areas including: Genome Biology & Genetics, Cell Biology, Ecology & Evolutionary Biology, or Microbiology. The doctoral program is designed to train students to apply sophisticated quantitative techniques to solving basic and applied problems in biology. Students in this program will attain substantially greater quantitative skills than in traditional doctoral programs in the biological sciences, providing them with a competitive advantage in business, industry, government, and academia.

The  University Catalog  contains all UTA admission requirements for the Department of Biology Ph.D. program.

How to Apply

First, read the general instructions for applying to the graduate program and download the checklists available on the Office of Graduate Studies webpage:  www.uta.edu/admissions/graduate/apply

International applicants have specific requirements, and a separate admissions checklist, available here: https://www.uta.edu/admissions/apply/international-graduate  

Additionally, the Department requests that applicants fill out the following questionnaire as part of the application package:  https://common.forms.uta.edu/view.php?id=2034157   Students are accepted into our Ph.D. program as either direct admit or rotation students. 

Direct admit: To qualify as a direct admit, an applicant must secure the backing of a faculty member before applying, mention their faculty support in their letter of intent, and ask that their faculty member reach out to Dr. Mark Pellegrino ( [email protected] ).

To secure the support of a faculty member before applying, begin by visiting our Biology faculty page: https://www.uta.edu/academics/schools-colleges/science/departments/biology/faculty - research/faculty , review the research focuses of the labs, and contact individual faculty members to discuss your research interests. Explain how your interests align with their lab and inquire if they would consider having you join their lab as a new Ph.D. candidate.

Rotation program: Applicants would choose this option if they are unsure which area of research interests them. Please indicate your interest in the rotation program in your letter of intent. Applicants admitted into the program will perform three rotations in three separate labs. At the end of the rotation period, applicants are placed in a home lab under the guidance of a Rotation Program Committee. Applicants will select their rotation labs only after being admitted into the Ph.D. program. Please note that applications to the rotation program are only considered for the Fall semester . For more information about the rotation program, please contact the chair of the Rotation Program Committee, Dr. Mark Pellegrino ( [email protected] ).

ALL APPLICATIONS ARE REVIEWED ON THE FOLLOWING SCHEDULE:

Questions? Please contact : [email protected]  

ADMISSION REQUIREMENTS

Students interested in pursuing a Ph.D. in the Department of Biology must apply to either the “B.S. to Ph.D. track” or the “Ph.D. program”, with the corresponding prerequisites:

B.S. to Ph.D. track:  A bachelor's degree in biology, or a bachelor's degree in some other discipline with at least 12 hours of advanced level coursework (junior or senior level courses) in biology. This is the Ph.D. track designed for students with a B.S and without master’s degree, which tends to be the majority of applicants.

Ph.D. program:  This track is for students that already have a master's degree in biology, or at least 30 hours of graduate level coursework in biology. A minimum undergraduate GPA of 3.0 (4.0 scale), as calculated by the Graduate School, is expected. An applicant's GPA in science courses will be taken into consideration. If an applicant has a master's degree, the GPA from their master's degree will also be considered. For applicants with up to 30 hours of graduate coursework but no master's degree, the GPA from those hours will be also be taken into consideration.

GRE:   The Graduate Record Exam (GRE) is no longer required for admission. 

Application, scores, and transcripts:   Your online application will include instructions for uploading transcripts, test scores, and any other required materials. 

International students:  Students whose native language is not English must provide a score on the Test of Spoken English (TSE) of at least 45, a minimum score of 23 on the Speaking portion of the TOEFL iBT exam or a minimum score of 7 on the Speaking portion of the IELTS exam. See the following website for complete details:  https://www.uta.edu/admissions/apply/international-graduate

Letters of recommendation:  Three letters  of recommendation are required for admission to the program. Ideally one or two letters should come from professors or instructors who can speak to your academic abilities; others may come from employers or other individuals who can comment on your work ethic and potential for graduate study. Letters should be emailed directly from the letter writer to [email protected] . If you have questions regarding letters of recommendation, please email [email protected] and include your UTA ID #, provided within your online application.

Letter of intent:  Please email a Letter of Intent to [email protected] and include the following information:

• Your full name and UTA ID #.
• Indicate that you are seeking the Ph.D. degree in Quantitative Biology
• Tell us which research track or types of research you are interested in. Examples include: Genome Biology & Genetics, Cell & Developmental Biology, Ecology & Evolutionary Biology, or Microbiology
• Provide names of any specific faculty member(s) that you are interested in working with if you have particular preferences. Note, however, that it is also OK if you are not sure what faculty member you are interested in working with (which is common), in which case, we will use your research interests above to help match you with potential faculty research mentors. We also encourage you to reach out to faculty members directly by email to discuss the potential for research in their lab, and if you have contacted any, please put this information in your letter of intent.
• How do you decide on which lab to conduct research in for your PhD? - Some students have a clear idea of their research interests, others may not. Both situations are common and our PhD program can account for both types of students, and students can decide on a single lab upfront to conduct research (direct admission) or decide to rotate in multiple labs during their first year, and decide which of these they will do their PhD work in in the second year. Our diverse department enables a broad range of research opportunities.
• Identify if you wish to be admitted directly to a single faculty member’s laboratory (and identify this faculty member), or indicate that you are interested in potentially rotating in multiple labs (2-3 labs) in the first year to make this decision. If interested in a lab rotation, please also indicate which faculty members’ labs are of interest, and if you have been in contact these faculty. More information about Rotations is provided below.
• Your Letter of Intent should also include details of your experience, training, any research experience you may have had, and motivations that you believe make you a strong candidate for success in a PhD program. You may also briefly discuss the long-term professional goals that motivate you to work towards a PhD. This is also an appropriate place to clarify details of your academic and work experience to date.

What is the letter of intent used for:

• The letter of intent is used to help the department link interested candidates with potential research professors/labs, and serves as an opportunity for the applicant to make a personal argument for why they believe they will be successful in conducting a PhD based on their experience and motivations. Your letter will be forwarded to the faculty members you indicate, but you are encouraged to also contact those faculty members directly (e.g., by email) to determine if they are accepting new students. It is therefore useful to also visit the listed  faculty pages  to find faculty research that you might be interested in and contact those faculty members.

PhD Funding:  Ph.D. students are guaranteed 5 years of financial support in the form of Graduate Teaching Assistantships (GTAs), funded by the department, or Graduate Research Assistantships (GRAs), funded by grants to individual faculty members. Many students are supported by a mixture of these two assistantships throughout their PhD. All Ph.D. students enrolled full-time and receiving an assistantship also receive reduced-cost in-state tuition regardless of residence, and a major portion of these tuition costs are supported by the assistantships.

Checking the status of your application:   You can check your application and admissions status through the MyMav student portal, where you can also review if you have any missing items needed to complete your application.   CHECK YOUR STATUS IN MYMAV . To confirm receipt of letters of recommendation, first check with your recommenders before contacting  [email protected] .  Please include your UTA ID #, available via MyMav, in any correspondence with the Biology Department.

Lab Rotations

The UTA Biology Department encompasses numerous laboratories that conduct research on a remarkably broad range of topics, and choosing a lab and research topic can be difficult. About half of entering students join a single lab as a direct admission. The other portion of students decide to rotate in multiple labs their first year. If several labs are of interest to you, consider signing up for lab rotations your first semester. Our lab rotation option allows you to experience multiple labs before making a final decision. However, please be sure to reach out to at least two potential rotation faculty to be sure they are able and interested in taking students for rotation. In your letter of intent (see  How To Apply  above), inform the Biology Department which labs are of interest to you (list up to three labs), and note which faculty have agreed to host you as a lab rotation student if you are accepted.

• If you decide to enter the PhD program as a rotation student, you can do ½ semester rotations in two host labs to get a sense of the research being conducted prior to making a final decision on which lab you will conduct your PhD research in.  
• For those students opting for a lab rotation, your letter of intent will be forwarded to the faculty members you indicate, but you should also contact those individuals directly to determine if they are taking new students and have the resources to support your research. During your first semester you will join/rotate through up to three labs in which you are interested and that are currently recruiting students. If no fit is found during your first semester, you have the option of rotating through additional labs in your second semester. Please visit the listed faculty pages to find faculty research that you might be interested in and contact those faculty members to set up a rotation. We feel confident that you will find an exciting and dynamic dissertation lab to join.  

BIOLOGY GRADUATE HANDBOOK [PDF]

The Biology Graduate Handbook is currently under revision. Up-to-date information regarding the PhD in Biology is available in the UTA Graduate Catalog . Questions regarding the Biology graduate program should be emailed to [email protected] .

• Cooperative Work Program
• Master's Program
• Graduate Program Handbook (PDF)

Location : Life Sciences Building, Room 337, 501 S. Nedderman Dr., Arlington, TX 76019 Mailing address : P.O. Box 19498 Phone : 817-272-2872

Social Media

Best Global Universities for Biology and Biochemistry

The best universities for biology and biochemistry show strength in producing research in the field and have top-notch reputations. This field includes many subjects, such as anatomy, physiology and pathology, which all share a core mission of analyzing the science of life. These are the world's top universities for biology and biochemistry. Read the methodology »

To unlock more data and access tools to help you get into your dream school, sign up for the  U.S. News College Compass !

Here are the best global universities for biology and biochemistry

Harvard university, massachusetts institute of technology (mit), stanford university, university of california berkeley, university of california san francisco, university of cambridge, university of california san diego, university of washington seattle, university of oxford, johns hopkins university.

See the full rankings

• Clear Filters

• # 1 in Best Universities for Biology and Biochemistry
• # 1 in Best Global Universities

Founded in 1636, Harvard University is the oldest higher education institution in the U.S. The bulk of Harvard's... Read More

• # 2 in Best Universities for Biology and Biochemistry
• # 2 in Best Global Universities

Massachusetts Institute of Technology, founded in 1861, is located in Cambridge, Massachusetts, near Boston. Around... Read More

• # 3 in Best Universities for Biology and Biochemistry
• # 3 in Best Global Universities

Stanford University was founded in 1885 and is located in California’s Bay Area, around 30 miles south of San Francisco... Read More

• # 4 in Best Universities for Biology and Biochemistry
• # 4 in Best Global Universities

The University of California—Berkeley is situated roughly 15 miles from San Francisco in what is known as the Bay Area... Read More

• # 5 in Best Universities for Biology and Biochemistry
• # 16 in Best Global Universities  (tie)

The University of California—San Francisco is a public institution that was founded in 1864. The health sciences-focused... Read More

• # 6 in Best Universities for Biology and Biochemistry
• # 8 in Best Global Universities

The University of Cambridge, located around 60 miles north of London, traces its history back to 1209. Around 19,000... Read More

• # 7 in Best Universities for Biology and Biochemistry
• # 20 in Best Global Universities

The University of California—San Diego is a public institution that was established in 1960. It is located in the La... Read More

• # 8 in Best Universities for Biology and Biochemistry
• # 6 in Best Global Universities

The University of Washington is a public institution that was founded in 1861. The school's oldest and largest campus in... Read More

• # 9 in Best Universities for Biology and Biochemistry
• # 5 in Best Global Universities

The exact date of the University of Oxford’s founding is unknown, but the school traces its roots back to at least 1096... Read More

• # 10 in Best Universities for Biology and Biochemistry
• # 10 in Best Global Universities

Johns Hopkins University is a private institution that was founded in 1876. The school has campuses located in and... Read More

Department of Biology

Study at one of the country's first biology departments.

The Johns Hopkins Department of Biology is home to 27 research laboratories investigating a wide range of biological problems, with the unifying goal of obtaining explanations in quantitative and molecular detail.

• Degrees Offered BA, BS, MS, PhD
• Major Biology, Molecular and Cellular Biology

Explore the Department

Undergraduate Studies

The Department of Biology offers a rigorous undergraduate and five-year combined master’s program. Students develop a detailed, nuanced view of biology; integrating knowledge at the molecular, cellular, and organismal levels.

Graduate Studies

The Program in Cell, Molecular, Developmental Biology, and Biophysics (CMDB) offers a doctoral degree.

JHU BioREU is a Research Experiences for Undergraduates Site funded by the National Science Foundation. The 10-week summer program provides an intensive, mentored research experience.

News & Announcements

Congratulations to Dr. Rejji Kuruvilla on Becoming the Vice Dean for Natural Sciences!

Dr. Rejji Kuruvilla, a professor in the Department of Biology, now also assumes the role of vice dean for natural sciences as of January 1, 2024, taking over from Bertrand […]

Richard E. McCarty, former Krieger School dean and renowned plant physiologist, dies at 85

An article featured in the HUB that can be found here, announced that Richard E. McCarty, died at home in Roland Park Place in Baltimore on February 23 due to […]

Disordered C-terminal domain drives spatiotemporal confinement of RNAPII to enhance search for chromatin targets

The Wu lab employed single-molecule tracking and spatiotemporal mapping in live yeast to demonstrate the necessity of the CTD in restricting RNAPII diffusion within a subnuclear area enriched with active […]

Biology Graduate Program Ranked in U.S. News & World Report

More than 30 University of Iowa colleges and programs were recognized among the best in their field in the U.S. News & World Report Best Graduate Schools rankings including Biology's graduate program! Click on the link above to see the full list. 

• Request Info
• Life at Dickinson
• Tuition & Aid
• News & Events
• Alumni in Action
• Dickinson Magazine
• Global Study and Engagement
• Sustainability
• Latest news /
• Biochemistry & Molecular Biology Major Jobs and Graduate School Acceptances 2024 /

Biochemistry & Molecular Biology Major Jobs and Graduate School Acceptances 2024

biochemistry & molecular biology

Claire Shearer

• UPMC Summer Rotational Internship, where I shadowed various medical specialties 

Charmi Patel

Sophia kovatsis.

How has Dickinson prepared you for life after graduation?

Working with Associate Professor of Biology Michael Roberts on the genetic mechanisms of acute myeloid leukemia has prepared me to better understand how cancer works on a molecular level. Student-faculty research at Dickinson has been a large part of my experience, allowing me to learn in a hands-on manner and connect with professionals in the field at the annual AACR meeting where we present our research.

Demi Gerovasilis

Saul torres nieto.

• Restorative Physical Therapy and Rehabilitation, Thorndale, Pa., physical therapy aide.
• University of Southern California, Los Angeles, Calif.
• INP at Dickinson for HEST certificate, intern. 

Dana Kleinman

• Louis Stokes Cleveland Veterans Affairs Medical Center, lab of Dr. Ivy Samuels, research assistant. 

Maritriny Galvez-Ceron

• UPMC Community General Osteopathic Hospital with Dr. Thomas Pineo, intern.

Taytum Robinson-Covert

• Department of Environmental Protection Borough of Laboratories (DEP BOL) in Harrisburg, Pa., organic chemistry intern.

Sarah Tepfer

Billy wilkerson.

• Hebert lab.
• MedNews Week, manager and codirector of research.

TAKE THE NEXT STEPS

• Request Information
• Read More Dickinson News

Published May 17, 2024

Events All events

Employee recognition brunch.

Graduate School

Biochemistry (ph.d.), biochemistry (ph.d.) | graduate.

Our Biochemistry doctoral students are at the forefront of biochemical research and molecular medicine, examining biological mechanisms underlying human disease; they are finding new ways to detect and attack diseases and immunological disorders like cancer, neurological disorders, and cardiovascular disease.

Graduates of the Ph.D. in Biochemistry program at Howard's Graduate School are prepared for careers at top research universities and senior-level research positions in biomedical and related industries. The program's key strengths in molecular microbiology, proteomics and genetics, bioinformatics, and drug design and discovery make us a nexus for collaborative investigations between biochemistry researchers and clinicians. You'll learn to apply biochemical techniques, including NMR spectroscopy, crystallography, and single-molecule methods as well as contemporary approaches to cell culture and genetic analysis to answer key questions about the pathogenesis of specific diseases and the development of effective drug therapies. You'll also enjoy the close mentorship of faculty who are committed to your professional development. Our faculty are experts in several areas of biochemistry, including analysis of molecular structure, proteomics and genetics, tumor biology, structural biology, enzymology, RNA catalysis, stress response, and RNA modification. As you advance in the program, you'll become increasingly involved in laboratory research and the critical analysis of biochemical literature. Our graduate seminar series offers a venue to present your early-stage research. Students may pursue a dual M.D./Ph.D. degree.

Program Snapshot

      ❱  72 credit hours        ❱  Full-time       ❱  On-campus format       ❱  Degree: Ph.D.       ❱  Dual degree: M.D./Ph.D.

Application Deadlines

Spring 2024 entry:         ❱  No spring entry 

Fall 2024 entry:         ❱  Dec. 1, 2023 (early deadline)       ❱  Feb. 15, 2024 (priority deadline)       ❱  Apr. 15, 2024 (final deadline)

Applicants should submit their applications as early as possible for earlier consideration of departmental funding opportunities. Applicants have until the final deadline to apply. However, applications will be reviewed on a rolling basis throughout the admissions cycle. 

Dr. Zaki Sherif

Dr. matthew george, jr., angela wilson, program details.

• Degree Classification: Graduate
• Related Degrees: M.D. / Ph.D., Ph.D.

Admission Requirements

Application for admission.

• Online GradCAS application
• Statement of purpose/ Statement of academic interest ( 500-1,000 words )
• GRE scores  not required
• Official transcripts sent to GradCAS
• 3 letters of recommendation
• Bachelor's degree from an accredited college or university or the international equivalent 
• Resume or Curriculum Vitae
• Autobiographical statement ( 500-750 words )

GRE Required?

Gre preferred minimums.

• GRE Verbal Reasoning: N/A
• GRE Quantitative Reasoning: N/A
• GRE Analytical Writing: N/A

GPA Required Minimums

• Overall GPA minimum: 3.0
• Undergrad GPA minimum: 3.0

Prerequisite Courses

The following course prerequisites are required. Applicants are required to have at least a B average in these prerequisites.   No expiration date for recommended prerequisites.

• Biology (college-level courses, 8 semester credit hrs)
• General Chemistry (college-level courses, 8 semester credit hrs)
• Organic Chemistry (college-level courses, 8 semester credit hrs)
• Elementary Physical Chemistry (college-level course and lab, 4 semester credit hrs)
• Physics (college-level courses, 8 semester credit hrs)
• Calculus  (college-level course, 3 semester credit hrs)

Reference Requirements

Evaluator type accepted:

• Professor (Required)
• Supervisor/Manager
• Other 

Evaluator type not accepted:

• Family Member

Art, language courses get an XR boost

Honoring Excellence: Alumni Awards 2024

Dr. Chen named joint-editor of journal

Biological sciences, applying for admission, ecology & environmental science, global sustainability, health sciences, and general tracks, for consideration, applicants must submit the following:.

• Intent to pursue a thesis or non-thesis version of your degree (if applicable),
• How you developed your interest in biology
• Career goals
• How pursuing a Central master’s in Biology will further your education or career
• Potential advisor from our faculty
• Two letters of recommendation from college instructors familiar with your scientific abilities. Three letters for Global Sustainability and General
• $50 application fee
• Applicants must have GPA of 3.00 or higher in all post-baccalaureate coursework

Wildlife & Conservation Biology Track

For consideration, applicants must submit the following:  .

• An essay describing experiences that led to your interest in wildlife conservation or conservation biology
• Two letters of recommendation from college instructors or supervisors
• General GRE
• General Ecology
• Population Genetics or Conservation Genetics
• Elementary Statistics
• A course in Animal or Plant Physiology
• Applicants must have a GPA of 3.00 in all post-baccalaureate course work

Official transcripts may be sent directly to the Graduate Recruitment & Admissions Office:

Central Connecticut State University Graduate Recruitment & Admissions Office 1615 Stanley Street Davidson Hall 116 New Britain, CT 06050

[email protected]

• Admission Requirements

Graduate Recruitment & Admissions

Multiple outstanding fully-funded PhD positions available in Biological Sciences at GenEvo in Germany

Offer Description

Thinking of doing a PhD at the interface of  Molecular & Evolutionary Biology ? 

The Research Training Group 2526  "Gene Regulation in Evolution“ (GenEvo) ,  which is funded by the DFG (German Research Foundation), is offering projects on all levels of organismic complexity, from changes in genes and proteins to the evolution of symbiotic and parasitic behavior. As a GenEvo PhD student, you will join a community of passionate scientists who are applying a broad spectrum of methods on model and non-model organisms.

GenEvo brings together scientists from evolutionary and molecular biology. It aims to solve a number of recent research questions and to train a new , interdisciplinary generation of PhD students in subjects that are rarely taught in combination : evolutionary inference, epigenetics, gene regulation, omics techniques and bioinformatics.

The PhD students get  training and guidance in both research fields  to ensure an optimal collaboration within the RTG for best results and personal development.

For detailed information about each PhD program, including specific requirements and other important details, we recommend visiting the following links:

1.  Mechanistic insights into the evolution of circadian gene regulation

2.  Co-option of cilia proteins in gene regulation

3.  Geometric epigenetic adaptation

4.  Epigenetics and sex chromosomes through the lens of evolution

5.  Sex- and caste determination during development in the termite Reticulitermes flavipes

6.  The role of small RNAs in transgenerational plasticity of a clonal plant upon herbivory

7.  Molecular manipulation of host phenotype via regulatory interference

8.  The role of epigenetic mechanisms in the regulation of division of labor in the ant Temnothorax longispinosus

9.  Gene regulation and evolution of dormant states in bees 

Job Information

• Organisation/Company:     PhD Programme "Gene Regulation in Evolution"
• Department:                       Institute of Molecular Biology
• Research Field:                  Biological sciences
• Researcher Profile:            First Stage Researcher (R1)
• Country:                             Germany
• Application Deadline:        15 Jul 2024 - 12:00 (Europe/Berlin)
• Type of Contract:               Temporary
• Job Status:                         Full-time
• Is the job funded through the EU Research Framework Programme?: Not funded by an EU programme
• Is the Job related to staff position within a Research Infrastructure?:    No

What we offer:

• Exciting, interdisciplinary projects in a  vividly international  environment, with English as our working language
• Advanced training  in scientific techniques and professional skills
• Access to state-of-the-art  Core Facilities  and their technical expertise
• 14 funded PhD positions  (employment contract)
• A lively community of34 PhD students supported by 28 Principle Investigators
• Collaboration with the  International PhD Programme (IPP)  at IMB with more than 200 PhD students from 40 different countries

Within the programme the  Faculty of Biology of Mainz University  (JGU) and the  Institute of Molecular Biology  (IMB) collaborate — both  modern research institutions  located on the bustling campus of Mainz University in Germany. With a population of 210,000, of which about 40,000 are students, the city of  Mainz  is  charming  and  open-minded and within easy reach of cosmopolitan  Frankfurt and its international airport , the Rhine valley region with its  castles, vineyards, and nature reserves , and the equally picturesque cities of Wiesbaden and Heidelberg.

Requirements:

Are you an ambitious, young scientist looking to  push the boundaries of research  while interacting with colleagues from  multiple disciplines and cultures ? Then joining GenEvo is your opportunity to  give  your scientific career a flying start!

All you need is:

• Master or equivalent
• Motivation to contribute to the forefront of science in molecular and evolutionary biology
• Interactive personality & good command of English
• 2 letters of reference

The deadline for applications is 15 July 2024.  Interviews will take place on 9-10 September 2024. Starting date will be 1 January 2025.

For more details on the projects offered and how to apply via our online form, please visit  https://www.genevo-rtg.de/application

Additional Information

• Website for additional job details:     https://www.genevo-rtg.de/application

Work Location(s)

• Organisation/Company:       PhD Programme "Gene Regulation in Evolution"
• Country:                               Germany 
• City:                                      Mainz
• Postal Code:                         55128       
• Street:                                   Ackermannweg 4
• Where to apply:                      https://www.genevo-rtg.de/application

E-Mail: [email protected]

Website: EURAXESS WORLDWIDE

• Boise State News
• Share on Facebook
• Share on Twitter
• Share on LinkedIn
• Share through Email
• Campus Life and Student Success
• Health, Science and Technology

Boise State triple major awarded graduate research fellowship

College of Arts and Sciences recent graduate Sevio Stanton has been selected for prestigious research and education advancement by the National Science Foundation (NSF) Graduate Research Fellowship Program.

The program awards outstanding students in STEM-based graduate programs for their meritorious research and promising outlook. The five-year fellowship provides three years of financial support inclusive of an annual stipend of $37,000 to advance their graduate-level research.

In addition to Stanton, Boise State’s 2024 awardees include College of Engineering student Sarah Pooley and the following three alumni: Andrew Russell (’21, mechanical engineering) now at the University of Washington; Audrey Parker (’22, materials science and engineering) at Massachusetts Institute of Technology; and Kyle Fisch (’21, chemistry and biochemistry) at the University of Colorado at Boulder.

Meet Sevio Stanton

Excelling in one major is challenging enough, but Stanton did well in three – he graduated from Boise State in May 2024 with a triple major in physics, chemistry and biology.

“[I was] stirred by the wonder of what connections and insights might become apparent to me if I were able to view the world through the combined lenses,” he said.

When his focus shifted toward physics, he retained the other two majors to evaluate chemistry and biology from an insider’s perspective.

Stanton applied the lessons learned in the classroom to practical lab work in physics and chemistry. He took research assistantships at Boise State, working with professors Brian Jackson, Ken Cornell, Brian McClain and Daniel Fologea. He also took prestigious internships at the Massachusetts Institute of Technology and Fermilab, studying particle physics and artificial intelligence applications in astronomy.

“My professors have all been very helpful by providing letters of recommendation for the undergraduate research programs that I have applied to,” Stanton said. “Dr. Fologea was a tremendous collaborative help with the research proposal for the National Science Foundation fellowship program.”

Stanton is heading to the University of Colorado at Boulder in the fall of 2024 where he’ll pursue a doctorate in physics. He’s particularly interested in high-energy theory, cosmology and dark matter. The NSF and Department of Energy’s Computational Science Graduate Fellowship programs will support his studies.

Office of Communications and Marketing