Ph.D. Program

The training for a Ph.D. in Biology is focused on helping students achieve their goals of being a successful research scientist and teacher, at the highest level. Students work closely with an established advisor and meet regularly with a committee of faculty members to facilitate their progress. The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. 

There are two tracks within the Biology Ph.D. program:

  • Cell, Molecular and Organismal Biology
  • Ecology and Evolution

(Previously a part of the Department of Biology Hopkins Marine Station is now a part of the Oceans Department within  Stanford Doerr School of Sustainability )

All  tracks are focused on excellence in research and teaching in their respective areas; where there are differences between the tracks, they are indicated in the links below. 

Requirements & Forms

Dissertation defense, cellular and molecular biology training program, stanford biology ph.d. preview program, career development resources.

Systems, Synthetic, and Quantitative Biology​

Share this page.

Harvard was one of the first institutions to offer a program to explore this exciting new field. The program’s core curriculum includes courses on the methods and logic that shape research, how to conceptualize and present research, and an introduction to the faculty’s research.

The program has 48 faculty located in the Faculty of Arts and Sciences, Harvard Medical School, and Harvard-affiliated teaching hospitals including Dana-Farber Cancer Institute, Mass General, and Boston Children’s Hospital. SSQB is one of 14 PhD programs in the Harvard Integrated Life Sciences program that collectively gives you access to over 900 faculty research groups situated in the heart of Boston’s biotech hub. Our students are working on projects that range from fundamental problems in biology to translational research, whose goal is to directly affect medicine and global sustainability.

Graduates of the program have gone on to faculty positions at prestigious institutions such as MIT and Princeton University, while others are now industry leaders as startup founders or as decision-makers at companies including Boston Consulting Group, Yumanity Therapeutics, McKinsey & Company, and Regeneron.

Additional information on the graduate program is available from the Systems, Synthetic, and Quantitative Biology PhD Program , and requirements for the degree are detailed in Policies .

Admissions Requirements

Please review admissions requirements and other information before applying. You can find degree program-specific admissions requirements below and access additional guidance on applying from the Systems, Synthetic, and Quantitative Biology PhD Program .

Academic Background

Applicants typically have a background in biology, physics, chemistry, computer science, engineering, or mathematics and work to forge a new approach to biology that combines theoretical and experimental approaches. The typical student has a strong background in one of the disciplines relevant to systems biology and an interest in interdisciplinary research.

Standardized Tests

GRE General: Optional 

Contacting Faculty

Applicants should indicate their faculty of interest in the application. You are not required to contact any faculty in advance but are welcome to.

Applications are reviewed by the admissions committee during December and early January. Selected applicants are notified if they have been chosen for an on-campus interview. These visits provide students with the opportunity to meet with faculty and current students and to get a better feel for our community and the types of research conducted here. Applicants invited for an interview who reside overseas and cannot visit the Harvard campus may interview remotely.

Theses & Dissertations

Theses & Dissertations for Systems, Synthetic, and Quantitative Biology​

See list of Systems, Synthetic, and Quantitative Biology​ faculty

APPLICATION DEADLINE

Questions about the program.

Molecular and Cell Biology

Doctoral Program

The Department of Molecular & Cell Biology at the University of California, Berkeley offers a Ph.D. program focused on the molecular mechanisms inherent to life. This program integrates research with a modern training curricula, teaching, and career mentorship. Our Department is highly interdisciplinary - comprising the Divisions of Cell Biology, Development & Physiology, Immunology and Molecular Medicine, Neurobiology, Biochemistry, Biophysics & Structural Biology, and Genetics, Genomics, and Development – and this is reflected in our students and training. The program is also highly collaborative with related programs and Institutes on campus, thus allowing students the flexibility to explore all aspects of modern biological research. Please click on the links below to learn more about our areas of research on the main department website or use the menu at the top to navigate to areas of interest within the graduate program.

Cell Biology, Development & Physiology

Immunology and molecular medicine, neurobiology, biochemistry, biophysics & structural biology, molecular therapeutics, genetics, genomics, evolution, and development.

  • Skip to Content
  • Catalog Home
  • Institution Home
  • Graduate Catalog /
  • School of Arts & Sciences /

Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2023 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

Sample Plan of Study

Print options.

Print this page.

The PDF will include all information unique to this page.

A PDF of the entire 2023-24 catalog.

A PDF of the 2023-24 Undergraduate catalog.

A PDF of the 2023-24 Graduate catalog.

Doctor of Philosophy (PhD) in Biology

The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of full-time study.

A summary of Biology PhD student expectations by year can be found here . Full details can be found in the Graduate Program Guide .

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program.

How to Apply    Frequently Asked Questions

Learning Outcomes

1. Demonstrate academic mastery in one of three areas of Biology: Ecology, Evolution, Behavior & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

2. Attain research expertise , including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

3. Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

4. Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Coursework Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits.  See the  Graduate School of Arts & Sciences (GRS) Transfer of Credits policy  for more details. 

  • 2 semesters of Progress in Research Seminars (2 credits each):

Cell & Molecular Biology:  BI 583 & BI 584 (CM section)

Ecology, Behavior, Evolution & Marine Biology:  BI 579 & BI 580

Neurobiology:   BI 583 & BI 584 (NEURO section)

  • 1 semester grant-writing course (2 credits):

Cell & Molecular Biology:  BI 581

Ecology, Behavior, Evolution & Marine Biology: BI 671

Neurobiology: BI 581

  • 1 semester pedagogy course (1 credit): All Biology PhD students take  BI 697
  • 1 quantitative course (3–4 credits): from a list of recommended courses .
  • Additional courses vary by specific track below:

Cell & Molecular Biology

1. GRS BI 791/GRS BI 792 Graduate Rotation Credits (2 credits each/4 credits total)

2. GRS BI 753 Advanced Molecular Biology  (4 credits)

3. GRS MB 721 Graduate Biochemistry  (4 credits)

4. GRS BI 735 Advanced Cell Biology  (4 credits)

5. Two electives (8 credits, 500-level and above), see the Graduate Program Guide for recommendations

6. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology

1. Six electives  (24 credits, 500-level and above), see the Graduate Program Guide for recommendations

2. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology PhD candidate coursework is highly variable. Students, in consultation with advisors, develop a plan of coursework and research. Students are required to take a minimum of 32 credits of coursework. The remainder of the credits should be research.

Neurobiology

2. GRS BI 755 Cellular and Systems Neuroscience  (4 credits)

3. GRS BI 741 Neural Systems: Functional Circuit Analysis  (4 credits)

4. Four electives  (16 credits, 500-level and above), see the Graduate Program Guide for recommendations

5. Research credits  (remaining credits)

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge . The course provides guidance and training on pedagogy and other aspects of graduate school.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the  Cell & Molecular Biology and Ecology, Behavior, Evolution & Marine Biology curricula , this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation & Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the  GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Graduation Timeline

Forms and additional information about PhD graduation can be found on the GRS website .

9 – 12 months before proposed graduation date

  • Dissertation Prospectus & approval form due to Graduate Program Specialist for departmental review and submission to GRS

Semester prior to your intended graduation cycle

  • Intent to Graduate Form completed online

About 2 months before dissertation defense

  • Meet with Graduate Program Specialist, Director of Graduate Studies, and Faculty Advisor to review requirements for defense
  • Arrange for Special Service Appointments if committee members are not BU faculty
  • Send first draft of dissertation to readers

Once defense date is confirmed with committee

  • Reserve room(s) for public seminar and defense

At least three weeks prior to dissertation defense

  • Schedule of Final Oral Exam with Abstract Approval due to GRS
  • Properly formatted draft of dissertation submitted as PDF to [email protected]

At least two weeks prior to dissertation defense

  • Send dissertation to all committee members

At least one week prior to dissertation defense

  • Send program information to Graduate Program Specialist

See the Graduate Program Guide for final dates to submit dissertation to ETD

  • Submit final dissertation to ETD (online submission)
  • Complete electronic signature page and forward confirmation to Graduate Program Specialist along with a copy of your submitted dissertation

MS Degree (En Route to PhD)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam), and has completed 32 credits of graduate-level coursework (not including research) may apply to the Graduate School for an MS degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two: A PhD student who has taken, but has not advanced to candidacy based on the PhD qualifying examination, may still receive an MS degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for a MS degree, and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Interdisciplinary Study Options

Biology PhD students have the option to participate in the Boston University Graduate Program in Urban Biogeoscience and Environmental Health (BU URBAN), the National Science Foundation Research Traineeship Program Understanding the Brain: Neurophotonics (NSF NRT UtB: Neurophotonics), and the Biogeoscience Advanced Graduate Certificate Program . These programs require separate applications in addition to the standard Biology PhD application; those interested in BU URBAN are encouraged pre-apply .

Time Limits

Officially, the PhD must be completed within seven years after the first registration for doctoral study. PhD degrees are conferred in either May, August, or January, as specified on the GRS website . In addition, the PhD candidacy expires after the fifth anniversary of passing the Qualifying Examination. Petitions to extend this deadline are possible at the discretion of the Director of Graduate Studies, the Department Chair, and the Dean of the Graduate School, and can be obtained from the Office of the Graduate School of Arts & Sciences.

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program. PhD students are encouraged to apply for fellowships and grants at funding agencies. All domestic students should apply for NSF Graduate Research Fellowships in the Fall semester of their first or second year. 

Travel Grants may be available to assist students in their travel to professional scientific meetings; students presenting papers or posters on their research will receive first consideration.

Common Types of Funding:

Dean’s Fellowships: These are non-service fellowships allocated to first-year PhD students that do not have immediate teaching requirements.

Teaching Fellowships: These provide a stipend plus full tuition and fees for up to four full courses per semester plus a 2-credit teaching course. Teaching responsibilities usually require approximately 20 hours per week. Full or partial awards may be given.

Doctoral Research Fellowships: These awards are given to students who assist individual faculty with specific areas of research. These Research Fellowships provide a stipend and full tuition. The supervising faculty member determines the specific duties of the Research Fellow.

In addition to the above funding sources, several competitive Department awards and fellowships are available to graduate students in the Department of Biology.

Back to Top

  • Program Finder
  • Admissions Services
  • Course Directory
  • Academic Calendar
  • Hybrid Campus
  • Lecture Series
  • Convocation
  • Strategy and Development
  • Implementation and Impact
  • Integrity and Oversight
  • In the School
  • In the Field
  • In Baltimore
  • Resources for Practitioners
  • Articles & News Releases
  • In The News
  • Statements & Announcements
  • At a Glance
  • Student Life
  • Strategic Priorities
  • Inclusion, Diversity, Anti-Racism, and Equity (IDARE)
  • What is Public Health?

Doctor of Philosophy (PhD)

Offered By: Department of Biochemistry and Molecular Biology

Onsite | Full-Time | 5 – 6 years

  • MAS Application Fee Waiver Requirements
  • Master of Arts (MA) in Geography and Environmental Engineering
  • Master of Arts and Master of Science in Public Health (MA/MSPH)
  • Master of Arts in Public Health Biology (MAPHB)
  • Master of Bioethics (MBE)
  • Mission, Vision, and Values
  • Student Experience
  • Program Outcomes
  • For Hopkins Undergraduate Students
  • Master of Health Science (MHS) - Department of Biochemistry and Molecular Biology
  • Master of Health Science (MHS) - Department of Epidemiology
  • Alumni Update
  • MHS Combined with a Certificate Program
  • Master of Health Science (MHS) - Department of Molecular Microbiology and Immunology
  • Alumni Highlights
  • Post-Baccalaureate Program in Environmental Health for Pre-Medicine Students
  • Bachelor's/MHS in Health Economics and Outcomes Research
  • MHS HEOR Careers
  • Frequently Asked Questions
  • Master of Health Science (MHS)
  • Concurrent School-Wide Master of Health Science Program in Biostatistics
  • Master of Health Science - Department of Population, Family and Reproductive Health
  • Master of Health Science Online (MHS) - Department of Population, Family and Reproductive Health
  • Careers in Health Economics
  • Core Competencies
  • Meet the Director
  • What is Health Economics
  • MPH Capstone Schedule
  • Concentrations
  • Online/Part-Time Format
  • Requirements

Tuition and Funding

  • Executive Board Faculty
  • Master of Science (MS) in Geography and Environmental Engineering
  • Independent Professional Project and Final Essay 
  • Program Objectives and Outcomes
  • Internships
  • Master of Science (ScM) - Department of Biochemistry and Molecular Biology
  • Master of Science (ScM) - Department of Biostatistics
  • Master of Science (ScM) - Department of Epidemiology
  • Master of Science (ScM) - Department of Molecular Microbiology and Immunology
  • ScM Faculty Advisers
  • Master of Science in Engineering (MSE) in Geography and Environmental Engineering
  • Bachelor's/MSPH in Health Policy
  • FAQ for MSPH in Health Policy
  • Field Placement Experience
  • MSPH Capstone
  • MSPH Practicum
  • Required and Elective Courses
  • Student Timeline
  • Career Opportunities
  • 38-Week Dietetics Practicum
  • Completion Requirements
  • MSPH/RD Program FAQ
  • Program Goals
  • Master's Essay Titles
  • Application Fee Waiver Requirements
  • Doctor of Philosophy (PhD) - Department of Biostatistics
  • Doctor of Philosophy (PhD) - Department of Epidemiology
  • Program Goals and Expectations
  • Doctor of Philosophy (PhD) - Department of Molecular Microbiology and Immunology
  • Doctor of Philosophy (PhD) - Department of Population, Family and Reproductive Health
  • Doctor of Philosophy (PhD) in Clinical Investigation
  • Track in Environmental Sustainability, Resilience, and Health
  • Track in Exposure Sciences and Environmental Epidemiology
  • Track in Health Security
  • Track in Toxicology, Physiology and Molecular Mechanisms
  • PhD in Geography and Environmental Engineering Faculty Advisers
  • Recent Graduates and Dissertation Titles
  • PhD Funding
  • PhD TA Requirement
  • Recent Dissertation Titles
  • JHU-Tsinghua Doctor of Public Health
  • Core Course Requirements
  • Concentration in Women’s and Reproductive Health
  • Custom Track
  • Concentration in Environmental Health
  • Concentration in Global Health: Policy and Evaluation
  • Concentration in Health Equity and Social Justice
  • Concentration in Health Policy and Management
  • Concentration in Implementation Science
  • Meet Current Students
  • Combined Bachelor's / Master's Programs
  • Concurrent MHS Option for BSPH Doctoral Students
  • Concurrent MSPH Option for JHSPH Doctoral students
  • Doctor of Medicine and Doctor of Philosophy (MD/PhD)
  • Adolescent Health Certificate Program
  • Bioethics Certificate Program
  • Climate and Health Certificate Program
  • Clinical Trials Certificate Program
  • Community- Based Public Health Certificate Program
  • Demographic Methods Certificate Program
  • Environmental and Occupational Health Certificate Program
  • Epidemiology for Public Health Professionals Certificate Program
  • Evaluation: International Health Programs Certificate Program
  • Food Systems, the Environment and Public Health Certificate Program
  • Frequently Asked Questions for Certificate Programs
  • Gender and Health Certificate Program
  • Gerontology Certificate Program
  • Global Digital Health Certificate Program
  • Global Health Certificate Program
  • Global Health Practice Certificate Program
  • Health Communication Certificate Program
  • Health Disparities and Health Inequality Certificate Program
  • Health Education Certificate Program
  • Health Finance and Management Certificate Program
  • Health and Human Rights Certificate Program
  • Healthcare Epidemiology and Infection Prevention and Control Certificate Program
  • Humane Sciences and Toxicology Policy Certificate Program
  • Humanitarian Health Certificate Program
  • Implementation Science and Research Practice Certificate Program
  • Injury and Violence Prevention Certificate Program
  • International Healthcare Management and Leadership Certificate Program
  • Leadership for Public Health and Healthcare Certificate Program
  • Lesbian, Gay, Bisexual, Transgender, and Queer (LGBTQ) Public Health Certificate Program
  • Maternal and Child Health Certificate Program
  • Mental Health Policy, Economics and Services Certificate Program
  • Non-Degree Students General Admissions Info
  • Pharmacoepidemiology and Drug Safety Certificate Program
  • Population Health Management Certificate Program
  • Population and Health Certificate Program
  • Product Stewardship for Sustainability Certificate Program
  • Public Health Advocacy Certificate Program
  • Public Health Economics Certificate Program
  • Public Health Informatics Certificate Program
  • Public Health Practice Certificate Program
  • Declaration of Intent - Public Health Preparedness
  • Public Health Training Certificate for American Indian Health Professionals
  • Public Mental Health Research Certificate Program
  • Quality, Patient Safety and Outcomes Research Certificate Program
  • Quantitative Methods in Public Health Certificate Program
  • Requirements for Successful Completion of a Certificate Program
  • Rigor, Reproducibility, and Responsibility in Scientific Practice Certificate Program
  • Risk Sciences and Public Policy Certificate Program
  • Spatial Analysis for Public Health Certificate Program
  • Training Certificate in Public Health
  • Tropical Medicine Certificate Program
  • Tuition for Certificate Programs
  • Vaccine Science and Policy Certificate Program
  • Online Student Experience
  • Online Programs for Applied Learning
  • Barcelona Information
  • Fall Institute Housing Accommodations
  • Participating Centers
  • Registration, Tuition, and Fees
  • Agency Scholarship Application
  • General Scholarship Application
  • UPF Scholarship Application
  • Course Evaluations
  • Online Courses
  • Registration
  • General Institute Tuition Information
  • International Students
  • Directions to the Bloomberg School
  • All Courses
  • Important Guidance for ONSITE Students
  • D.C. Courses
  • Registration and Fees
  • Cancellation and Closure Policies
  • Application Procedures
  • Career Search
  • Current Activities
  • Current Trainees
  • Related Links
  • Process for Appointing Postdoctoral Fellows
  • Message from the Director
  • Program Details
  • Admissions FAQ
  • Current Residents
  • Elective Opportunities for Visiting Trainees
  • What is Occupational and Environmental Medicine?
  • Admissions Info
  • Graduates by Year
  • Compensation and Benefits
  • How to Apply
  • Academic Committee
  • Course Details and Registration
  • Tuition and Fees
  • ONLINE SOCI PROGRAM
  • Principal Faculty
  • Johns Hopkins RAPID Psychological First Aid
  • General Application
  • JHHS Application
  • Areas of Study
  • Important Dates
  • Our Faculty
  • Welcome Letter
  • Descripción los Cursos
  • Programa en Epidemiología para Gestores de Salud, Basado en Internet
  • Consultants
  • Britt Dahlberg, PhD
  • Joke Bradt, PhD, MT-BC
  • Mark R. Luborsky, PhD
  • Marsha Wittink, PhD
  • Rebekka Lee, ScD
  • Su Yeon Lee-Tauler, PhD
  • Theresa Hoeft, PhD
  • Vicki L. Plano Clark, PhD
  • Program Retreat
  • Mixed Methods Applications: Illustrations
  • Announcements
  • 2023 Call for Applications
  • Jennifer I Manuel, PhD, MSW
  • Joke Bradt, PhD
  • Josiemer Mattei, PhD, MPH
  • Justin Sanders, MD, MSc
  • Linda Charmaran, PhD
  • Nao Hagiwara, PhD
  • Nynikka R. A. Palmer, DrPH, MPH
  • Olayinka O. Shiyanbola, BPharm, PhD
  • Sarah Ronis, MD, MPH
  • Susan D. Brown, PhD
  • Tara Lagu, MD, MPH
  • Theresa Hoft, PhD
  • Wynne E. Norton, PhD
  • Yvonne Mensa-Wilmot, PhD, MPH
  • A. Susana Ramírez, PhD, MPH
  • Animesh Sabnis, MD, MSHS
  • Autumn Kieber-Emmons, MD, MPH
  • Benjamin Han, MD, MPH
  • Brooke A. Levandowski, PhD, MPA
  • Camille R. Quinn, PhD, AM, LCSW
  • Justine Wu, MD, MPH
  • Kelly Aschbrenner, PhD
  • Kim N. Danforth, ScD, MPH
  • Loreto Leiva, PhD
  • Marie Brault, PhD
  • Mary E. Cooley, PhD, RN, FAAN
  • Meganne K. Masko, PhD, MT-BC/L
  • PhuongThao D. Le, PhD, MPH
  • Rebecca Lobb, ScD, MPH
  • Allegra R. Gordon, ScD MPH
  • Anita Misra-Hebert, MD MPH FACP
  • Arden M. Morris, MD, MPH
  • Caroline Silva, PhD
  • Danielle Davidov, PhD
  • Hans Oh, PhD
  • J. Nicholas Dionne-Odom, PhD RN ACHPN
  • Jacqueline Mogle, PhD
  • Jammie Hopkins, DrPH, MS
  • Joe Glass, PhD MSW
  • Karen Whiteman, PhD MSW
  • Katie Schultz, PhD MSW
  • Rose Molina, MD
  • Uriyoán Colón-Ramos, ScD MPA
  • Andrew Riley, PhD
  • Byron J. Powell, PhD, LCSW
  • Carrie Nieman MD, MPH
  • Charles R. Rogers, PhD, MPH, MS, CHES®
  • Emily E. Haroz, PhD
  • Jennifer Tsui, Ph.D., M.P.H.
  • Jessica Magidson, PhD
  • Katherine Sanchez, PhD, LCSW
  • Kelly Doran, MD, MHS
  • Kiara Alvarez, PhD
  • LaPrincess C. Brewer, MD, MPH
  • Melissa Radey, PhD, MA, MSSW
  • Sophia L. Johnson, PharmD, MPH, PhD
  • Supriya Gupta Mohile, MD, MS
  • Virginia McKay, PhD
  • Andrew Cohen, MD, PhD
  • Angela Chen, PhD, PMHNP-BC, RN
  • Christopher Salas-Wright, PhD, MSW
  • Eliza Park MD, MS
  • Jaime M. Hughes, PhD, MPH, MSW
  • Johanne Eliacin, PhD, HSPP
  • Lingrui Liu ScD MS
  • Meaghan Kennedy, MD
  • Nicole Stadnick, PhD, MPH
  • Paula Aristizabal, MD
  • Radhika Sundararajan, MD
  • Sara Mamo, AuD, PhD
  • Tullika Garg, MD MPH FACS
  • Allison Magnuson, DO
  • Ariel Williamson PhD, DBSM
  • Benita Bamgbade, PharmD, PhD
  • Christopher Woodrell MD
  • Hung-Jui (Ray) Tan, MD, MSHPM
  • Jasmine Abrams, PhD
  • Jose Alejandro Rauh-Hain, MD
  • Karen Flórez, DrPH, MPH
  • Lavanya Vasudevan, PhD, MPH, CPH
  • Maria Garcia, MD, MPH
  • Robert Brady, PhD
  • Saria Hassan, MD
  • Scherezade Mama, DrPH
  • Yuan Lu, ScD
  • 2021 Scholars
  • Sign Up for Our Email List
  • Workforce Training
  • Cells-to-Society Courses
  • Course/Section Numbers Explained
  • Pathway Program with Goucher College
  • The George G. Graham Lecture

About the PhD in Biochemistry and Molecular Biology Program

In the Biochemistry and Molecular Biology PhD program, faculty, and students work together to increase knowledge of the biochemical and molecular bases of normal and abnormal cellular processes. Our program trains students to be successful independent scientists and gives them the knowledge, research training, and leadership skills to continue to provide new insights into the biomedical issues that have a profound impact on public health.

Students engage in a rigorous course curriculum and a range of structured and informal activities outside the classroom and lab to build their skills. They will pursue their thesis research in the lab of one of our over forty training faculty across the Johns Hopkins Bloomberg School of Public Health and the Johns Hopkins School of Medicine.

Visit our dedicated PhD program website to learn more about the diverse research training opportunities of the program.

PhD in Biochemistry and Molecular Biology Program Highlights

Our position within the School of Public Health provides a unique setting in which students learn how biochemistry, molecular biology, physical chemistry, cell biology, and genetics can be used to solve significant problems in public health and medicine. Our program offers:

  • Training faculty from across the School of Public Health and the School of Medicine
  • A strong grounding in the science of biomedical and public health research through a core curriculum that includes courses taught by leading experts from the Schools of Public Health and Medicine
  • Training outside the lab and classroom in key skills such as communications and leadership
  • Opportunities to build strong communications skills through a range of speaking venues including journal club, research colloquium, department retreats, and national meetings
  • Access to the Johns Hopkins School of Medicine Professional Development and Career Office , offering excellent career services and professional development,  including the BMB-required OPTIONS program, a guided process of career exploration for paths from medicine to biotech to academia and beyond for careers paths from medicine to biotech to academia and beyond
  • Opportunities to participate in community service and outreach, with a focus on our East Baltimore neighborhoods, through the Johns Hopkins University community engagement and service-learning center, SOURCE

Training faculty across the School of Public Health and the School of Medicine

Schools that students can take courses in: Public Health, Arts & Sciences, Medicine, and Engineering

Two-month rotations in the first year prior to selecting thesis lab

Average number of incoming students in the BMB PhD degree program each year

What Can You Do With a PhD In Biochemistry And Molecular Biology?

The Biochemistry and Molecular Biology PhD program prepares students for a range of biomedical and health sciences careers, including in academia, industry, policy, and beyond. Visit the Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries.

Sample Careers

  • Research Scientist
  • Science Policy Adviser
  • Biotech Executive
  • Senior Scientist
  • Patent Lawyer
  • Science Policy Analyst/Advocate
  • Science Writer/Journalist
  • Biological Sciences Teacher

Topic Areas

The BMB PhD program faculty conduct research to gain new insights into the cellular and molecular mechanisms underlying normal and abnormal cellular processes, and their relevance as targets for improving health and treating disease. Our training program places particular emphasis on mechanistic approaches to research problems.

Common topic areas within our faculty's diverse research interests include:

  • Biophysics and Structural Biology
  • Cancer Biology
  • Chemical Biology and Proteomics
  • Cell Biology
  • Cellular Stress and Cell Signaling
  • Genetics, Genomics, and Gene Regulation
  • Immunology and Infectious Diseases
  • Translational Research

Curriculum for the PhD in Biochemistry and Molecular Biology

The BMB PhD offers students a rigorous course curriculum, including a set of common core classes from the Schools of Public Health and Medicine. A rich array of seminar programs and journal clubs are also available to all students.

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Courses in core curriculum

Minimum elective credits

Seminars on current research presented by experts from across Johns Hopkins and other biomedical research institutions

Courses available across Johns Hopkins Schools of Public Health, Medicine, and Arts and Sciences

Admissions Requirements

For the general admissions requirements see our How to Apply page. The specific program also requires:

Prior Work Experience

Laboratory research experience (from academia, industry, etc.) is required

Prior Coursework

Strong background in the sciences, particularly in chemistry, biochemistry, or biology

Standardized Test Scores

Standardized test scores (GRE) are optional for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Program Faculty Spotlight

Ashani Weeraratna

Ashani T. Weeraratna

Ashi Weeraratna, PhD, studies how cancer cells move to distant sites and how changes in the normal cells around a tumor contribute to their movement, especially as we age.

best phd programs for biology

Michael J. Matunis

Michael Matunis, PhD, studies how protein modification by SUMO—the small ubiquitin-related modifier—drives changes in key cellular pathways from stress response to DNA repair.

Jennifer Kavran

Jennifer M. Kavran

Jennifer Kavran, PhD, MS, MPhil, is a biophysicist who investigates how cells communicate with each other and their environment.

best phd programs for biology

Danfeng Cai

Danfeng Cai, PhD, combines advanced microscopy, genomics, and proteomics to tease out the functions of protein condensates in cells, with a focus on cancer.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1 , 202 3 . VTSI applicants are eligible for an application fee waiver , but the fee waiver must be requested by November 15, 202 3 and prior to submission of the SOPHAS application.

Vivien Thomas

All full-time PhD students receive the following support for all years of the program: full tuition and fees, individual health insurance, University Health Services fee, vision insurance, dental insurance, and a stipend for living expenses for students who remain in good academic standing. PhD students are required to serve as a teaching assistant for at least one term, in either their 2nd or 3rd year.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU starting in Fall 2023 or beyond can apply to receive a $1500 need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Mike Matunis, PhD PhD Program Director

Roza Selimyan , PhD BMB Executive Director for Academic Affairs and Education Programs

Erika Vaitekunas Administrative Specialist

[email protected]

Compare Programs

  • Check out similar programs at the Bloomberg School to find the best fit.
  • Master of Science (ScM), Offered by BMB
  • Master of Health Science (MHS), Offered by BMB
  • Master of Health Science (MHS), Offered by MMI
  • Doctor of Philosophy (PhD), Offered by MMI
  • Master of Health Science (MHS) in Environmental Health
  • Doctor of Philosophy (PhD) in Environmental Health

Are you seeking one-on-one college counseling and/or essay support? Limited spots are now available. Click here to learn more.

50 Best Colleges for Biology – 2024

May 8, 2024

best biology colleges

Biological sciences is the most popular major for those planning to one day apply to med schools. Many biology majors will also continue their education beyond a bachelor’s pursuing Master’s and Ph.D. degrees. With all of these challenges ahead, you’ll want to choose a renowned institution that is one of the best biology colleges and will help you emerge from four years of study ready to compete in a competitive job or graduate school market. The colleges and universities that made our list of best colleges for biology are known for producing knowledgeable graduates who have had ample opportunity for hands-on learning in research laboratories.

When choosing a degree program, you will likely have the opportunity to concentrate in a specific area such as Computational Biology and Mathematical Biology, Ecology and Evolutionary Biology, Mechanisms of Disease, Molecular and Cell Biology, and Neurobiology.

Methodology 

Click here to read our methodology for the Best Colleges for Biology.

Best Colleges for Biology

Here’s a quick preview of the first biology institutions that made our list. Detailed profiles and stats can be found when you scroll below.

1) Swarthmore College

2) University of California, Los Angeles

3) Harvard University

4) Massachusetts Institute of Technology

5) California Institute of Technology

6) Williams College

7) Stanford University

8) Johns Hopkins University

9) University of Pennsylvania

10) Cornell University

All of the schools profiled below have stellar reputations in the field of biology and commit substantial resources to undergraduate education. For each of the best biology colleges, College Transitions will provide you with—when available—the university’s:

  • Cost of Attendance
  • Acceptance Rate
  • Median  SAT
  • Median  ACT
  • Retention Rate
  • Graduation Rate

We will also include a longer write-up of each college’s:

  • Academic Highlights – Includes facts like student-to-faculty ratio, average class size, number of majors offered, and most popular majors.
  • Professional Outcomes – Includes info on the rate of positive outcomes, companies employing alumni, and graduate school acceptances.

Swarthmore College

Swarthmore College

  • Swarthmore, PA

Academic Highlights: Swarthmore offers forty undergraduate programs and runs 600+ courses each academic year. Small, seminar-style courses are the norm—an outstanding 33% of sections enroll fewer than ten students, and 70% contain a maximum of nineteen students. Social science degrees are the most commonly conferred, accounting for 24% of all 2022 graduates. Future businessmen/women, engineers, and techies are also well-positioned, given Swat’s incredibly strong offerings in economics, engineering, and computer science.

Professional Outcomes: 68% of Class of 2022 grads entered the workforce shortly after graduation. Popular industries included education (17%), consulting (16%), and financial services (13%); the median starting salary was $60,000. Google is a leading employer of Swarthmore grads followed by Amazon, Goldman Sachs, IBM, and a number of the top universities.  18% of 2022 grads pursued advanced degrees, with 35% pursuing a PhD, 35% entering master’s programs, 10% heading to law school, and 7% matriculating into medical school.

  • Enrollment: 1,625
  • Cost of Attendance: $81,376
  • Median SAT: 1500
  • Median ACT: 33
  • Acceptance Rate: 7%
  • Retention Rate: 96%
  • Graduation Rate: 94%

University of California, Los Angeles

University of California, Los Angeles

  • Los Angeles, CA

Academic Highlights: UCLA offers 125 majors in 100+ academic departments, and more than 60 majors require a capstone experience that results in the creation of a tangible product under the mentorship of faculty members. The most commonly conferred degrees are in the social sciences (25%), biology (16%), psychology (11%), mathematics (8%), and engineering (7%). Departmental rankings are high across the board, especially in computer science, engineering, film, fine and performing arts, mathematics, and political science.

Professional Outcomes: UCLA grads flow most heavily into the research, finance, computer science, and engineering sectors. High numbers of recent grads can be found at Disney, Google, EY, Teach for America, Amazon, and Oracle. Hundreds also can be found at Bloomberg, Deloitte, Mattel, Oracle, and SpaceX. The average starting salary exceeds $55,000. 16% of recent grads enrolled directly in a graduate/professional school, with other CA-based institutions like Stanford, Pepperdine, USC, Berkeley, and Loyola Marymount being the most popular.

  • Enrollment: 33,040 (undergraduate); 15,010 (graduate)
  • Cost of Attendance: $38,517 (in-state); $71,091 (out-of-state)
  • Median SAT: Test Blind
  • Median ACT: Test Blind
  • Acceptance Rate: 9%
  • Retention Rate: 97%
  • Graduation Rate: 93%

Harvard University

Harvard University

  • Cambridge, MA

Academic Highlights: There are 50 undergraduate fields of study referred to as concentrations; many are interdisciplinary. Even with a graduate population of over 14,000 to cater to, undergraduate class sizes still tend to be small, with 42% of sections having single-digit enrollments and 71% being capped at nineteen. Economics, government, and computer science are the three most popular areas of concentration at Harvard. Biology, chemistry, physics, math, statistics, sociology, history, English, and psychology all sit atop most departmental ranking lists.

Professional Outcomes: The Crimson Class of 2022 saw 15% of students head directly into graduate/professional school. Of the graduates entering the world of work (virtually everyone else), 58% were entering either the consulting, finance, or technology field. Over 1,000 Harvard alumni presently work for Google and over 500 for Microsoft, McKinsey & Company, and Goldman Sachs. Turning our attention to those moving on to graduate school, Harvard grads with at least a 3.5 GPA typically enjoy acceptance rates into medical school of 90% or greater.

  • Enrollment: 7,240
  • Cost of Attendance: $79,450
  • Median SAT: 1550
  • Median ACT: 35
  • Acceptance Rate: 3%
  • Retention Rate: 98%
  • Graduation Rate: 98%

Massachusetts Institute of Technology

Massachusetts Institute of Technology

Academic Highlights: Undergraduates pursue one of 57 majors and 59 minors at this world-class research institution that continues to be one of the world’s most magnetic destinations for math and science geniuses.  The student-to-faculty ratio is an astonishing 3-to-1, and over two-fifths of all class sections have single-digit enrollments, and 70% of courses contain fewer than twenty students. The highest numbers of degrees conferred in 2022 were in the following majors: engineering (31%), computer science and engineering (28%), mathematics (10%), and the physical sciences (7%).

Professional Outcomes: The Class of 2023 saw 29% of its members enter the world of employment and 43% continue on their educational paths. The top employers included Accenture, Amazon, Microsoft, Goldman Sachs, Google, General Motors, the US Navy, Apple, Bain & Company, and McKinsey. The mean starting salary for an MIT bachelor’s degree holder was $95,000. The most frequently attended graduate schools are a who’s who of elite institutions including MIT itself, Stanford, Caltech, Harvard, and the University of Oxford.

  • Enrollment: 4,657
  • Cost of Attendance: $82,730
  • Acceptance Rate: 4%
  • Retention Rate: 99%
  • Graduation Rate: 95%

California Institute of Technology

California Institute of Technology

  • Pasadena, CA

Academic Highlights: Across all divisions, there are 28 distinct majors. Possessing an absurdly favorable 3:1 student-to-faculty ratio, plenty of individualized attention is up for grabs. Class sizes are not quite as tiny as the student-to-faculty ratio might suggest, but 70% of courses enroll fewer than twenty students, and 28% enroll fewer than ten. Computer science is the most popular major, accounting for 38% of all degrees conferred. Engineering (30%), the physical sciences (20%), and mathematics (6%) also have strong representation.

Professional Outcomes: Caltech is a rare school that sees six-figure average starting salaries for its graduates; in 2022, the median figure was $120,000. Forty-three percent of recent grads went directly into the workforce and found homes at tech giants such as Google, Intel, Microsoft, Apple, and Meta. A healthy 46% of those receiving their diplomas in 2022 continued directly on the higher education path, immediately entering graduate school. Ninety-seven percent of these students were admitted to one of their top-choice schools.

  • Enrollment: 982
  • Cost of Attendance: $86,886

Williams College

Williams College

  • Williamstown, MA

Academic Highlights: The school’s 25 academic departments offer 36 majors and a number of concentrations rather than minors. An unparalleled 40% of courses have fewer than ten students enrolled; the median class size is 12 students. Programs in economics, English, history, math, and political science are especially renowned, and the greatest number of degrees are conferred in the social sciences (26%), the physical sciences (10%), math and statistics (9%), psychology (9%), and computer science (7%).

Professional Outcomes: Among the Class of 2022, 92% were employed or continuing their educational journey within six months of graduating. Business and education typically attract the most students, with popular companies/organizations including Apple, Google, Goldman Sachs, The New York Times Co., the Peace Corps, and Teach for America. The median annual income for 2022 grads was $75,000. 75% pursue an advanced degree within five years of leaving Williams, with the most frequently attended graduate programs being Harvard, Columbia, and Yale.

  • Enrollment: 2,152 (undergraduate); 53 (graduate)
  • Cost of Attendance: $81,160
  • Median SAT: 1520
  • Acceptance Rate: 8%

Stanford University

Stanford University

  • Palo Alto, CA

Academic Highlights: Stanford has three undergraduate schools: the School of Humanities & Sciences, the School of Engineering, and the School of Earth, Energy, and Environmental Sciences. 69% of classes have fewer than twenty students, and 34% have a single-digit enrollment. Programs in engineering, computer science, physics, mathematics, international relations, and economics are arguably the best anywhere. In terms of sheer volume, the greatest number of degrees are conferred in the social sciences (17%), computer science (16%), engineering (15%), and interdisciplinary studies (13%).

Professional Outcomes: Stanford grads entering the working world flock to three major industries in equal distribution: business/finance/consulting/retail (19%); computer, IT (19%); and public policy and service, international affairs (19%). Among the companies employing the largest number of recent grads are Accenture, Apple, Bain, Cisco, Meta, Goldman Sachs, Google, McKinsey, Microsoft, and SpaceX. Other companies that employ hundreds of Cardinal alums include LinkedIn, Salesforce, and Airbnb. Starting salaries for Stanford grads are among the highest in the country.

  • Enrollment: 8,049 (undergraduate); 10,236 (graduate)
  • Cost of Attendance: $87,833
  • Median SAT: 1540

Johns Hopkins University

Johns Hopkins University

  • Baltimore, MD

Academic Highlights: With 53 majors as well as 51 minors, JHU excels in everything from its bread-and-butter medical-related majors to international relations and dance. Boasting an enviable 6:1 student-to-faculty ratio and with 78% of course sections possessing an enrollment under 20, face time with professors is a reality. Many departments carry a high level of clout, including biomedical engineering, chemistry, English, and international studies. Biology, neuroscience, and computer science, which happen to be the three most popular majors, can also be found at the top of the national rankings.

Professional Outcomes: The Class of 2022 saw 94% of graduates successfully land at their next destination within six months of exiting the university; 66% of graduates entered the world of employment and a robust 19% went directly to graduate/professional school. The median starting salary across all majors was $80,000 for the Class of 2022. JHU itself is the most popular choice for graduate school. The next most frequently attended institutions included Columbia, Harvard, Yale, and MIT.

  • Enrollment: 6,044
  • Cost of Attendance: $86,065

University of Pennsylvania

University of Pennsylvania

  • Philadelphia, PA

Academic Highlights : 90 distinct degrees are available across four schools: the College of Arts & Sciences, the College of Applied Science and Engineering, the College of Nursing, and the world-renowned Wharton School. The greatest number of students pursue degrees in business (19%), social sciences (14%), biology (11%), health sciences (9%), engineering (9%), and computer science (9%). The university boasts an exceptional 26% of courses with an enrollment under ten and 59% with an enrollment under twenty as well as multiple ways for undergrads to conduct research.

Professional Outcomes: 75% of Class of 2022 grads were employed within six months of graduating, and 18% were in graduate school. Finance attracted the highest percentage of grads (30%) followed by consulting (20%), technology (15%), and healthcare (10%). Employers hiring the greatest number of 2022 grads included JPMorgan, Boston Consulting Group, McKinsey, Bain & Company, Meta, and Goldman Sachs. The median starting salary for all graduates is $80,000. For those continuing their educational journeys, the most popular move is to remain at Penn, followed by Columbia and Harvard.

  • Enrollment: 9,760 (undergraduate); 13,614 (graduate)
  • Cost of Attendance: $89,028
  • Graduation Rate: 96%

Cornell University

Cornell University

Academic Highlights: A diverse array of academic programs includes 80 majors and 120 minors spread across the university’s seven schools/colleges. Classes are a bit larger at Cornell than at many other elite institutions. Still, 55% of sections have fewer than 20 students. Most degrees conferred in 2022 were in computer science (17%), engineering (13%), business (13%), and biology (13%). The SC Johnson College of Business houses two undergraduate schools, both of which have phenomenal reputations.

Professional Outcomes: Breaking down the graduates of the College of Arts and Sciences, the largest school at Cornell, 68% entered the workforce, 28% entered graduate school, 1% pursued other endeavors such as travel or volunteer work, and the remaining 3% were still seeking employment six months after receiving their diplomas. The top sectors attracting campus-wide graduateswere financial services (18%), technology (17%), consulting (15%), and education (10%). Of the students from A&S going on to graduate school, 15% were pursuing JDs, 5% MDs, and 22% PhDs.

  • Enrollment: 15,735
  • Cost of Attendance: $88,150
  • Median ACT: 34

Amherst College

Amherst College

  • Amherst, MA

Academic Highlights: A 7:1 student-to-faculty ratio allows for 66% of courses to have fewer than twenty students and 32% to have single-digit enrollments. By senior year, 98% of seniors report feeling close enough to a faculty member to ask for a letter of recommendation. Amherst possesses strong offerings across the board, most notably in economics, English, history, mathematics, and law The social sciences account for 22% of degrees conferred, while 14% are in mathematics, 11% in biology, and 7% in computer science

Professional Outcomes: Six months after graduation, 93% of the Class of 2022 had already found its way into the world of employment, graduate school, or a volunteer organization.  The largest employers of Amherst grads includes Google, Deloitte, Morgan Stanley, and Goldman Sachs. The schools where the highest number of Amherst grads can be found pursuing advanced degrees include MIT, Dartmouth, and the University of Pennsylvania. Fifty to sixty Amherst grads apply to medical school each year, and the acceptance rate hovers around 75-80%.

  • Enrollment: 1,898
  • Cost of Attendance: $84,840
  • Retention Rate: 95%
  • Graduation Rate: 92%

University of Chicago

University of Chicago

  • Chicago, IL

Academic Highlights: There are 53 majors at UChicago, but close to half of all degrees conferred are in four majors: economics, biology, mathematics, and political science, all of which have particularly sterling reputations. Economics alone is the selection of roughly one-fifth of the undergraduate population. Over 75% of undergrad sections have an enrollment of nineteen or fewer students, and undergraduate research opportunities are ubiquitous as 80% of students end up working in a research capacity alongside a faculty member.

Professional Outcomes: On commencement day, 99% of the Class of 2023 were employed or continuing their education. Business and financial services (30%) and STEM (12%) were the two sectors that scooped up the most graduates, but public policy and consulting were also well-represented. The most popular employers of recent grads include Google, JPMorgan, Goldman Sachs, McKinsey & Company, Bank of America, Citi, and Accenture. For those heading to grad school, the top seven destinations are Yale, Columbia, Penn, MIT, Stanford, UCLA, and Johns Hopkins.

  • Enrollment: 7,653 (undergraduate); 10,870 (graduate)
  • Cost of Attendance: $89,040
  • Acceptance Rate: 5%

Duke University

Duke University

Academic Highlights: The academic offerings at Duke include 53 majors, 52 minors, and 23 interdisciplinary certificates. Class sizes are on the small side—71% are nineteen or fewer, and almost one-quarter are less than ten. A stellar 5:1 student-to-faculty ratio helps keep classes so reasonable even while catering to five figures worth of graduate students. Computer Science is the most popular area of concentration (11%), followed by economics (10%), public policy (9%), biology (8%), and computer engineering (7%).

Professional Outcomes: At graduation, approximately 70% of Duke diploma-earners enter the world of work, 20% continue into graduate schools, and 2% start their own businesses. The industries that attract the largest percentage of Blue Devils are tech (21%), finance (15%), business (15%), healthcare (9%), and science/research (6%). Of the 20% headed into graduate school, a hefty 22% are attending medical school, 18% are in PhD programs, and 12% are entering law school. The med school acceptance rate is 85%, more than twice the national average.

  • Enrollment: 6,640
  • Cost of Attendance: $85,238
  • SAT Range: 1490-1570
  • ACT Range: 34-35
  • Acceptance Rate: 6%
  • Graduation Rate: 97%

Haverford College

Haverford College

  • Haverford, PA

Academic Highlights: Haverford offers 31 majors, 32 minors, 12 concentrations, and eleven consortium programs—areas of study that can be pursued at partner campuses. The school’s 9:1 student-to-faculty ratio and exclusive emphasis on undergraduate education lead to exceptionally intimate classes, 33% of which have fewer than 10 students, and 72% have fewer than 20. The most popular areas of study at Haverford include the social sciences (24%), biology (14%), psychology (11%), physical sciences (10%), computer science (9%), and mathematics (7%).

Professional Outcomes: Six months after leaving Haverford, 63% of the Class of 2022 had found employment, 19% had enrolled in graduate school, and 9% were still job hunting. Employers hiring multiple recent Haverford grads include Epic, JP Morgan Chase Bank, Boston Consulting Group, Goldman Sachs, the National Institutes of Health, and the Children’s Hospital of Philadelphia. Of the 19% of 2022 grads who elected to continue their education, the most commonly entered fields of study were STEM (51%) and medicine/health (15%).

  • Enrollment: 1,421
  • Cost of Attendance: $87,180
  • Median SAT: 1490
  • Acceptance Rate: 14%
  • Graduation Rate: 91%

Princeton University

Princeton University

  • Princeton, NJ

Academic Highlights: 39 majors are available at Princeton. Just under three-quarters of class sections have an enrollment of 19 or fewer students, and 31% have fewer than ten students. Princeton is known for its commitment to undergraduate teaching, and students consistently rate professors as accessible and helpful. The Engineering Department is widely recognized as one of the country’s best, as is the School of Public and International Affairs.

Professional Highlights: Over 95% of a typical Tiger class finds their next destination within six months of graduating. Large numbers of recent grads flock to the fields of business and engineering, health/science, & tech. Companies presently employing hundreds of Tiger alumni include Google, Goldman Sachs, Microsoft, McKinsey & Company, Morgan Stanley, IBM, and Meta. The average salary ranges from $40k (education, health care, or social services) to $100k (computer/mathematical positions). Between 15-20% of graduating Tigers head directly to graduate/professional school.

  • Enrollment: 5,604 (undergraduate); 3,238 (graduate)
  • Cost of Attendance: $86,700

University of California, San Diego

University of California, San Diego

  • San Diego, CA

Academic Highlights: There are 140+ undergraduate majors offered at UCSD, and all students join one of eight undergraduate colleges meant to forge flourishing communities within the larger university. Biology has the highest representation of all majors (19%) followed by engineering (12%), the social sciences (11%), and computer science (9%). UCSD’s computer science and engineering programs have stellar reputations in the corporate and tech communities, and programs in biology, economics, and political science are among the best anywhere.

Professional Outcomes: Employers of recent graduates included the Walt Disney Company, Tesla, NBC Universal, PwC, Northrop Grumman, and EY. More than 1,000 current Google employees are UC San Diego alumni, and Qualcomm, Amazon, and Apple all employ 500+ each. The median early career salary is $65,000 across all majors, placing the university in the top 10 public universities in the country. UCSD also fares well in measures of its return-on-investment potential.

  • Enrollment: 33,096 (undergraduate); 8,386 (graduate)
  • Cost of Attendance: $31,830 (in-state); $64,404 (out-of-state)
  • Acceptance Rate: 25%
  • Retention Rate: 93%
  • Graduation Rate: 88%

Columbia University

Columbia University

  • New York, NY

Academic Highlights: Columbia offers 100+ unique areas of undergraduate study as well as a number of pre-professional and accelerated graduate programs.  Class sizes at Columbia are reasonably small and the student-to-faculty ratio is favorable; however, in 2022, it was revealed that the university had been submitting faulty data in this area. It is presently believed that 58% of undergraduate courses enroll 19 or fewer students. The greatest number of degrees are conferred in the social sciences (22%), computer science (15%), engineering (14%), and biology (7%).

Professional Outcomes: Examining the most recent graduates from Columbia College and the Fu Foundation School of Engineering & Applied Science, 73% had found employment within six months, and 20% had entered graduate school. The median starting salary for graduates of Columbia College/Columbia Engineering is above $80,000. Many graduates get hired by the likes of Amazon, Goldman Sachs, Morgan Stanley, Google, Citi, McKinsey, and Microsoft.

  • Enrollment: 8,832
  • Cost of Attendance: $89,587

Bowdoin College

Bowdoin College

  • Brunswick, ME

Academic Highlights: Class sizes are small—64% contain fewer than twenty students—and 21% have fewer than ten students. The student-faculty ratio is 9:1. More than half of Bowdoin undergrads report interacting with a professor outside of regular class time at least once per week. The greatest percentage of degrees are conferred in the social sciences (30%), biology (13%), area/ethnic/gender studies (8%), computer science (7%), and mathematics (7%). Economics and government and legal studies are two of the more popular majors within the social sciences.

Professional Outcomes: An examination of three recent years’ worth of outcomes data reveals that one year after graduation, between 73 and 77% of recent grads have found full-time employment, and 15% have gone directly into graduate school. Of those entering graduate school, 48% were enrolled in master’s programs, 23% in PhD programs, 13% in law school, and 8% in med school. The top twenty graduate schools attended, by volume, in the last five years make an exclusive list including six Ivies along with Duke, MIT, Johns Hopkins, and Stanford.

  • Enrollment: 1,915
  • Cost of Attendance: $82,600
  • Median SAT: 1510

University of North Carolina at Chapel Hill

University of North Carolina at Chapel Hill

  • Chapel Hill, NC

Academic Highlights: Undergraduates can choose from 74 bachelor’s degree programs in a number of schools and colleges, the largest of which is the College of Arts & Sciences. 44% of classes have a student enrollment under 20. The social sciences (15%), biology (12%), media/journalism (9%), computer science (8%), and business (6%) are the areas in which the most degrees are conferred. The Kenan-Flager Business School is internationally renowned and requires separate admission. Other strong programs include those in chemistry, journalism, psychology, and political science.

Professional Outcomes: Six months after leaving Chapel Hill, 97% of 2022 grads had entered employment, military service, or graduate school. Among the for-profit companies that hire the most graduates are Wells Fargo, IBM, Cisco, Deloitte, EY, Google, Microsoft, Amazon, Oracle, McKinsey & Company, and Goldman Sachs. In the nonprofit sector, a large number of alumni are employed by AmeriCorps, NIH, Teach for America, and the Peace Corps. The average starting salary is $70,619. 18% of 2022 grads enrolled directly in graduate/professional school.

  • Enrollment: 20,210 (undergraduate); 11,739 (graduate)
  • Cost of Attendance: $27,036 (in-state); $60,040 (out-of-state)
  • Median SAT: 1450
  • Median ACT: 32
  • Acceptance Rate: 17%

University of California, Berkeley

University of California, Berkeley

  • Berkeley, CA

Academic Highlights: More than 150 undergraduate majors and minors are available across six schools: the College of Letters and Science, the College of Chemistry, the College of Engineering, the College of Environmental Design, the College of Natural Resources, and the Haas School of Business. Many departments have top international reputations including computer science, engineering, chemistry, English, psychology, and economics. 22% of sections contain nine or fewer students, and over 55% of students assist faculty with a research project or complete a research methods course.

Professional Outcomes: Upon graduating, 49% of Cal’s Class of 2022 had already secured employment, and 20% were headed to graduate school. Business is the most popular sector, attracting 62% of employed grads; next up are industrial (17%), education (8%), and nonprofit work (7%). The median starting salary was $86,459 across all majors. Thousands of alumni can be found in the offices of Google, Apple, and Meta, and 500+ Golden Bears are currently employed by Oracle, Amazon, and Microsoft. The school is the number one all-time producer of Peace Corps volunteers.

  • Enrollment: 32,831 (undergraduate); 12,914 (graduate)
  • Cost of Attendance: $48,574 (in-state); $82,774 (out-of-state)
  • Acceptance Rate: 11%

Yale University

Yale University

  • New Haven, CT

Academic Highlights: Yale offers 80 majors, most of which require a one- to two-semester senior capstone experience. Undergraduate research is a staple, and over 70% of classes—of which there are over 2,000 to choose from—have an enrollment of fewer than 20 students, making Yale a perfect environment for teaching and learning. Among the top departments are biology, economics, global affairs, engineering, history, and computer science. The social sciences (26%), biology (11%), mathematics (8%), and computer science (8%) are the most popular areas of concentration.

Professional Outcomes: Shortly after graduating, 73% of the Yale Class of 2022 had entered the world of employment and 18% matriculated into graduate programs. Hundreds of Yale alums can be found at each of the world’s top companies including Google, Goldman Sachs, McKinsey & Company, Morgan Stanley, and Microsoft. The most common industries entered by the newly hired were finance (20%), research/education (16%), technology (14%), and consulting (12%). The mean starting salary for last year’s grads was $81,769 ($120k for CS majors). Nearly one-fifth of students immediately pursue graduate school.

  • Enrollment: 6,590 (undergraduate); 5,344 (graduate)
  • Cost of Attendance: $87,705

Brown University

Brown University

  • Providence, RI

Academic Highlights: Students must choose one of 80+ “concentration programs,” but there are no required courses. Class sizes tend to be small—68% have fewer than twenty students—and 35% are comprised of nine or fewer students. Biology, economics, computer science, mathematics, and engineering are among the most popular areas of concentration at Brown; however, it is hard to distinguish any one program, because Brown possesses outstanding offerings across so many disciplines.

Professional Outcomes: Soon after receiving their Brown diplomas, 69% of graduates enter the world of employment. Companies employing the greatest number of Brown alums include Google, Microsoft, Goldman Sachs, Amazon, Morgan Stanley, Apple, McKinsey & Company, and Bain & Company. The Class of 2022 saw 27% of graduates go directly into graduate/professional school. Right out of undergrad, Brown students boasted an exceptional 81% admission rate to med school and an 81% admission rate to law school.

  • Enrollment: 7,639
  • Cost of Attendance: $84,828
  • Median SAT: 1530

Dartmouth College

Dartmouth College

  • Hanover, NH

Academic Highlights: Dartmouth sports 60+ majors and a stunning breadth of course selections for an institution of its size. The learning environment at Dartmouth is extraordinarily intimate. Not only do 61% of course sections have under twenty students, but 18% have single-digit enrollments. The student-to-faculty ratio is an outstanding 7:1. Top programs offered by Big Green include biology, economics, neuroscience, and government. The social sciences are the most popular, accounting for 32% of degrees conferred, followed by computer science (10%), mathematics (9%), engineering (9%), and biology (7%).

Professional Outcomes: A great reputation along with a passionate alumni network that is 80,000 strong leads Dartmouth grads to successful transitions into graduate school and the world of work. Included in the top ten employers of Dartmouth grads are a number of investment banks including Goldman Sachs, Morgan Stanley, Bain & Company, Citibank, and Deutsche Bank. Right off the bat, 52% of graduates make more than $70,000 in salary. Those pursuing graduate degrees often flock to the likes of Harvard, Columbia, and Princeton.

  • Enrollment: 4,458
  • Cost of Attendance: $87,793

Pomona College

Pomona College

  • Claremont, CA

Academic Highlights: There are 48 majors and minors to select from with the most popular being social sciences (23%), biology (13%), and computer science (12%). Majors in economics, international relations, chemistry, and mathematics receive especially high marks. More than 600 courses are on the menu at Pomona alone, but students can access any of the Claremont Consortium’s 2,700 courses. Pomona’s 8:1 student-to-teacher ratio leads to an average class size of only 15 students, and over 50% of the undergraduate population conduct research alongside a faculty member.

Professional Outcomes: 71% of the Class of 2022 were employed within six months of graduating. Overall, the largest number of alumni can be found at Google, Kaiser Permanente, Microsoft, Amazon, and Meta. Recently, economics degree-earners have landed jobs at Goldman Sachs, Wells Fargo, Morgan Stanley, or Accenture. Majors in the hard sciences frequently landed at top research laboratories and hospitals. Of the 21% of 2022 grads who were accepted directly into graduate school, the most frequently attended institutions included the University of Cambridge, Duke, Harvard, Caltech, UChicago, and Stanford.

  • Enrollment: 1,761
  • Cost of Attendance: $88,296

Carleton College

Carleton College

  • Northfield, MN

Academic Highlights: Students work closely with their professors, and the college is routinely rated atop lists of best undergraduate teaching institutions. Small classes are the norm with the average being only sixteen students. It offers 33 majors, the most popular of which are within the disciplines of the social sciences (19%), the physical sciences (14%), biology (11%), computer science (11%), mathematics (10%), and psychology (8%).

Professional Outcomes: Target, Epic Systems, Google, Wells Fargo, and Amazon all employ large numbers of graduates. Carleton is a breeding ground for future scholars as a ridiculously high number of graduates go on to earn PhDs. In fact, by percentage, Carleton is one of the top five producers in the country of future PhDs. They produce an incredible number of doctoral degree holders in the areas of economics, math, political science, sociology, chemistry, physics, biology, and history.

  • Enrollment: 2,034
  • Cost of Attendance: $82,167

Davidson College

Davidson College

  • Davidson, NC

Academic Highlights: With its small size, the impressive part of the college is the exceptional quality of its offerings, not the breadth of them, as only 37 majors are available. The student-to-faculty ratio is 9:1, which allows the college to ensure that 62% have fewer than twenty students and 24% have enrollments you can count on two hands. Overall, the average number of students per class is only 18. Top programs at Davidson include psychology, political science, chemistry, and English; biology is also quite popular, accounting for 12% of degrees conferred in 2022.

Professional Outcomes: Looking at the outcomes data for 2022 grads, 70% landed jobs within six months of graduation, 26% were enrolled in a graduate program, and 3% were still seeking employment. Of those who attended grad school, the highest number were in healthcare-related programs (including MDs), law school, and laboratory sciences. Significant numbers of students pursue advanced degrees at other Southern gems including Vanderbilt, Emory, Duke, Wake Forest, and UNC.

  • Enrollment: 1,927
  • Cost of Attendance: $76,450
  • Median SAT: 1430
  • Retention Rate: 94%

Northwestern University

Northwestern University

  • Evanston, IL

Academic Highlights : Northwestern is home to six undergraduate schools, including Medill, which is widely regarded as one of the country’s best journalism schools. The McCormick School of Engineering also achieves top rankings, along with programs in economics, social policy, and theatre. The social sciences account for the greatest number of degrees conferred (19%), followed by communications/journalism (13%), and engineering (11%). 45% of classes have nine or fewer students enrolled; 78% have fewer than twenty enrollees. 57% of recent grads had the chance to conduct undergraduate research.

Professional Outcomes: Six months after graduating, 69% of the Class of 2022 had found employment and 27% were in graduate school. The four most popular professional fields were consulting (18%), engineering (18%), business/finance (16%), and communications/marketing/media (13%). Employers included the BBC, NBC News, The Washington Post , NPR, Boeing, Google, IBM, Deloitte, PepsiCo, Northrop Grumman, and Goldman Sachs. Across all majors, the average starting salary was $73k. Of those headed straight to graduate school, engineering, medicine, and business were the three most popular areas of concentration.

  • Enrollment: 8,659 (undergraduate); 14,073 (graduate)
  • Cost of Attendance: $91,290

Emory University

Emory University

  • Atlanta, GA

Academic Highlights: This midsize university offers a diverse array of majors (80+) and minors (60+), and 30% of Emory students pursue more than one area of study. Over half of Emory’s student body works directly with a faculty member on academic research and 58% of courses have class sizes of under twenty students. Ultimately, the greatest number of students go on to earn degrees in the social sciences (15%), biology (14%), business (14%), health professions (12%), and mathematics (9%).

Professional Outcomes: Shortly after graduation, 66% of 2022 grads were already employed, and 96% had arrived at their next destination. The top employers of recent Emory grads include Deloitte, Epic, ScribeAmerica, Meta, Morgan Stanley, and Cloudmed. Graduates of the Goizueta Business School found strong starting salaries with an average of $81k.  In the last few years, multiple Emory grads/alums received acceptance letters from the following top law schools like Columbia, Berkeley, and Georgetown. Med school acceptances included Duke, Johns Hopkins, and Vanderbilt.

  • Enrollment: 7,101
  • Cost of Attendance: $83,702
  • Graduation Rate: 90%

Grinnell College

Grinnell College

  • Grinnell, IA

Academic Highlights: Grinnell offers 42 areas of study and over 500 courses each semester. Thanks to a 9:1 student/faculty ratio and no competition from graduate students, 64% of classes have fewer than twenty students, and roughly one-quarter of sections have single-digit student enrollments. A top producer of PhDs in both the sciences and the social sciences, Grinnell has strong offerings in biology, chemistry, physics, economics, and history. The largest number of degrees are conferred in the social sciences, foreign languages, biology, and computer science.

Professional Outcomes: Upon graduation, 62% of Grinnell students head into the workforce, 26% to graduate school, 2% to postgraduate service, and 2% earned fellowships. Multiple recent grads accepted positions with JPMorgan, Epic, and Google. The mean salary of those employed was roughly $64,397. Close to 40% of those entering graduate school pursue PhDs or master’s degrees in the hard sciences, 8% head to law school, and 3% were bound for medical school; 88% of all applicants were accepted into their first or second choice program.

  • Enrollment: 1,759
  • Cost of Attendance: $82,890
  • Median SAT: 1460

University of California, Santa Barbara

University of California, Santa Barbara

  • Santa Barbara, CA

Academic Highlights: There are 90 undergraduate majors across three schools: the College of Letters and Science, the College of Engineering, and the College of Creative Studies. The social sciences are the most popular area of study, accounting for 27% of the total degrees conferred. Biology (10%), math (9%), and psychology (9%) are next in popularity. The school has highly regarded programs in communication, computer science, engineering, physics, environmental science, and the performing arts. More than half of sections contain fewer than 20 students, and 72% enroll 29 or fewer.

Professional Outcomes: Within six months of earning their diplomas, 84% of grads had found employment. The most popular industries were science/research (16%), engineering/computer programming (14%), business (13%), finance/accounting (11%), and sales (10%). Top employers of recent grads include Google, EY, KPMG, Oracle, Amazon, IBM, and Adobe. Many alumni also can be found at Apple, Meta, Microsoft, and Salesforce. Two years after graduating, UCSB alumni make an average salary of $55k; more than half make $100k by mid-career.

  • Enrollment: 23,460 (undergraduate); 2,961 (graduate)
  • Cost of Attendance: $41,289 (in-state); $73,863 (out-of-state)
  • Acceptance Rate: 28%
  • Retention Rate: 92%
  • Graduation Rate: 86%

University of Notre Dame

University of Notre Dame

  • Notre Dame, IN

Academic Highlights: 75 majors are offered across six undergraduate colleges: the School of Architecture, the College of Arts and Letters, the Mendoza School of Business (one of the country’s best business schools), the College of Engineering, the Keough School of Global Affairs, and the College of Science. In 2022, the most degrees were conferred in business (20%), the social sciences (18%), engineering (12%), and biology (8%). A solid 60% of courses enroll fewer than 20 students, and 15% have single-digit numbers. 75% of Notre Dame undergrads study abroad.

Professional Outcomes: 69% of 2022 grads directly entered the world of employment, with the most common industries being financial services (21%), consulting (17%), technology (12%), and health services (9%). Massive numbers of alumni can be found at Deloitte, EY, PwC, IBM, Accenture, Booz Allen Hamilton, Google, Microsoft, Amazon, Goldman Sachs, JPMorgan, and McKinsey & Co. The median early-career salary was $76,000. Of the 20% of grads who went directly into their graduate/professional studies, 18% were pursuing medical degrees and 9% were studying law.

  • Enrollment: 8,971 (undergraduate); 4,134 (graduate)
  • Cost of Attendance: $86,125
  • Acceptance Rate: 13%

Rice University

Rice University

  • Houston, TX

Academic Highlights : Rice offers more than 50 majors across six broad disciplines: engineering, architecture, music, social science, humanities, and natural science. Programs in biology, biochemistry, cognitive science, and music are incredibly strong, while the School of Architecture and the George R. Brown School of Engineering are among the highest-ranking schools in their disciplines. One-third of computer science majors are female, almost twice the national average. Class sizes are ideally small with 66% containing fewer than 20 students and a median class size of only fourteen.

Professional Outcomes: Six months after graduation, 88% of Rice grads have found careers or a graduate school home. Companies that employ many recent grads include Deloitte, Capital One, JP Morgan Chase, Google, and Microsoft. Over one hundred alumni are also current employees of companies such as Shell, ExxonMobil, Chevron, Amazon, Accenture, and Meta. Across all majors, the average starting salary is $73k. One-third of graduates move directly into graduate or professional school, with Harvard, Yale, Stanford, MIT, Columbia, and Berkeley being the most popular destinations.

  • Enrollment: 4,494 (undergraduate); 4,178 (graduate)
  • Cost of Attendance: $78,278
  • Median SAT: 35

Washington University in St. Louis

Washington University in St. Louis

  • St. Louis, MO

Academic Highlights : WashU admits students into five schools, many of which offer nationally recognized programs: Arts & Sciences, the Olin School of Business, the School of Engineering & Applied Sciences, and the Art of Architecture programs housed within the Sam Fox School of Design and Visual Arts. The most commonly conferred degrees are in engineering (13%), social sciences (13%), business (13%), biology (11%), and psychology (10%). 66% of classes have fewer than 20 students, and over one-quarter have single-digit enrollments. 65% double major or pursue a minor.

Professional Outcomes: The Class of 2022 sent 52% of grads into the workforce and 28% into graduate and professional schools. Companies employing the highest number of WashU grads feature sought-after employers such as Amazon, Bain, Boeing, Deloitte, Google, IBM, Goldman Sachs, and Microsoft. Of the employed members of the Class of 2022 who reported their starting salaries, 79% made more than $60k. The universities welcoming the largest number of Bears included the prestigious institutions of Caltech, Columbia, Harvard, Penn, Princeton, and Stanford.

  • Enrollment: 8,132 (undergraduate); 8,880 (graduate)
  • Cost of Attendance: $83,760

University of Washington – Seattle

University of Washington – Seattle

  • Seattle, WA

Academic Highlights: 180+ undergraduate majors are offered across thirteen colleges/schools. Personal connections with professors abound as 55% of grads complete a faculty-mentored research project. The College of Engineering, which includes the College of Computer Science & Engineering, is one of the best in the nation; UW also boasts strong programs in everything from business to social work to environmental science. The most popular degrees are the social sciences (13%), biology (12%), computer science (11%), and business (8%).

Professional Outcomes: Within months of graduation, 73% of Class of 2022 grads were employed and 17% were continuing their education. The most popular employers of the Class of 2022 included Google, Amazon, Microsoft, Boeing, and KPMG. Across all living alumni, 6,000+ work for Microsoft, and 4000+ work for each of Boeing and Amazon. Of those headed to graduate/professional school, just over half remain in state, mostly at UW itself. Large numbers of 2022 grads also headed to Columbia, Johns Hopkins, and USC.

  • Enrollment: 36,872 (undergraduate); 16,211 (graduate)
  • Cost of Attendance: $34,554 (in-state); $63,906 (out-of-state)
  • Median SAT: 1420
  • Acceptance Rate: 48%
  • Graduation Rate: 84%

University of Virginia

University of Virginia

  • Charlottesville, VA

Academic Highlights: Undergrads can study within one of seven colleges/schools, which all offer many small classes; 15% boast single-digit enrollment and 48% contain 19 or fewer students. The McIntire School of Commerce and the School of Engineering and Applied Science have glowing reputations. Other notable strengths include computer science, economics, and political philosophy, policy, and law. The most popular degree areas are liberal arts/general studies (22%), the social sciences (14%), engineering (11%), business/marketing (8%), and biology (7%).

Professional Outcomes:  Upon receiving their degree, 95% of the Class of 2022 immediately joined the workforce–with an average starting salary of $90k–or headed directly to graduate school. The most popular industries were internet & software, higher education, and management consulting. Capital One (85), Deloitte (46), Amazon (38), and Bain & Co. (26) scooped up the greatest number of 2022 grads. UVA itself was the most popular grad school destination followed by Columbia, Virginia Commonwealth University, and Johns Hopkins.

  • Enrollment: 17,496 (undergraduate); 8,653 (graduate)
  • Cost of Attendance: $39,494-49,874 (in-state); $78,214-90,378 (out-of-state)
  • Median SAT: 1470
  • Acceptance Rate: 19%

University of Wisconsin – Madison

University of Wisconsin – Madison

  • Madison, WI

Academic Highlights: There are 230+ undergraduate majors offered across eight schools and colleges, including the top-ranked School of Business and College of Engineering as well as the College of Letters and Science, the College of Agricultural and Life Sciences, and the Schools of Nursing, Education, Pharmacy, and Human Ecology. Undergrads can expect a mix of large and small classes, with 44% of sections enrolling fewer than 20 students. Business (18%), biology (12%), the social sciences (11%), and engineering (10%) are most popular.

Professional Outcomes: In a recent year, 46% of job-seeking grads graduated with an offer.  Top employers included UW-Madison, Epic, Kohl’s, Oracle, Deloitte, and UW Health. Across all graduating years, companies employing 250+ alumni include Google, Target, Microsoft, Amazon, Apple, PwC, Accenture, and Meta. 28% of recent grads enrolled directly in graduate/professional school; the majority stayed at UW–Madison while others headed to Columbia, Northwestern, and Carnegie Mellon. The university is the top producer of Peace Corps volunteers.

  • Enrollment: 37,230 (undergraduate); 12,656 (graduate)
  • Cost of Attendance: $28,916 (in-state); $58,912 (out-of-state)
  • Median SAT: 1440
  • Median ACT: 30
  • Acceptance Rate: 49%
  • Graduation Rate: 89%

University of Michigan

University of Michigan

  • Ann Arbor, MI

Academic Highlights: There are 280+ undergraduate degree programs across fourteen schools and colleges, and the College of Literature, Science, and the Arts (LSA) enrolls the majority of students. The Ross School of Business offers highly rated programs in entrepreneurship, management, accounting, and finance. The College of Engineering is also one of the best in the country. By degrees conferred, engineering (15%), computer science (14%), and the social sciences (11%) are most popular. A solid 56% of classes have fewer than 20 students.

Professional Outcomes: Within three months of graduating, 89% of LSA grads are employed full-time or in graduate school, with healthcare, education, law, banking, research, nonprofit work, and consulting being the most popular sectors. Within three months, 99% of Ross grads are employed with a median salary of $90k. Top employers include Goldman Sachs, Deutsche Bank, EY, Morgan Stanley, PwC, Deloitte, and Amazon.  Within six months, 96% of engineering grads are employed (average salary of $84k) or in grad school. General Motors, Ford, Google, Microsoft, Apple, and Meta employ the greatest number of alumni.

  • Enrollment: 32,695 (undergraduate); 18,530 (graduate)
  • Cost of Attendance: $35,450 (in-state); $76,294 (out-of-state)
  • Acceptance Rate: 18%

University of California, Davis

University of California, Davis

Academic Highlights: UC Davis offers 100+ undergraduate majors across four schools: the College of Agricultural and Environmental Sciences, the College of Biological Sciences, the College of Engineering, and the College of Letters and Science. 50% engage in some type of research/creative project outside the classroom. The areas of study with the largest number of degrees awarded were biology, the social sciences, psychology, and engineering. Programs in engineering, computer science, and animal science are nationally renowned.

Professional Outcomes: Many recent grads found homes at Silicon Valley or other California-based employers. Corporations employing 200 or more Aggies include Genentech, Google, Apple, Cisco, Meta, Oracle, Amazon, Microsoft, Salesforce, and LinkedIn. Ten years out of school, median earnings rise to $112k. Within one year of graduating, 39% of Aggies elect to continue their education; the most popular degrees pursued are master’s, MDs or other health doctorates, law, and MBA/MPA.

  • Enrollment: 31,797 (undergraduate); 9,053 (graduate)
  • Cost of Attendance: $41,389 (in-state); $73,963 (out-of-state)
  • Acceptance Rate: 42%
  • Graduation Rate: 87%

Tufts University

Tufts University

  • Medford, MA

Academic Highlights: Three schools serve Tufts’ undergraduate population: the College of Arts & Sciences, the College of Engineering, and the School of the Museum of Fine Arts. The three schools combined offer more than 90 majors and minors; approximately one-third of all students double major, and half declare a minor. 15% of all courses see fewer than ten students enrolled, and 60% have sub-twenty enrollments. The most popular majors include international relations, economics, computer science, political science, and biology—all of which receive very high marks.

Professional Outcomes: Six months after earning their diplomas, 97% of 2022 graduates were employed or attending graduate school. The most commonly entered fields were finance, consulting, real estate (23%); engineering and technology (22%); health, life sciences, environmental (21%); and education, advocacy, social services (11%). Prolific employers of Tufts alums include Booz Allen Hamilton, JPMorgan, MITRE, Google, Deloitte, Amazon, Raytheon, Morgan Stanley, and BlackRock. Of the 21% of 2022 grads who went directly to graduate school, 85% were accepted into their first-choice institution.

  • Enrollment: 6,815 (undergraduate); 6,616 (graduate)
  • Cost of Attendance: $88,300
  • Acceptance Rate: 10%

Macalester College

Macalester College

  • St. Paul, MN

Academic Highlights: Students can choose from roughly 40 majors and over 800 courses that are offered each academic year . Being an undergraduate-only institution, Macalester students enjoy the full benefits of the school’s 10:1 student-to-faculty ratio. The average class size is only 17 students, and 14% of class sections have single-digit enrollments. Macalester possesses strong offerings across many different disciplines. Programs in economics, international studies, and mathematics are among the best anywhere.

Professional Outcomes: Six months after graduating, 95% of the Macalester Class of 2022 had found employment, graduate school, or a fellowship. Employers of recent grads include ABC News, Google, Goldman Sachs, Dow Chemical Company, McKinsey & Company, the ACLU, the National Cancer Institute, and National Geographic . Across all sectors, the average starting salary for recent grads was above $62k. Sixty percent of Mac grads pursue an advanced degree within six years of earning their bachelor’s.

  • Enrollment: 2,175
  • Cost of Attendance: $79,890
  • Retention Rate: 88%

Vanderbilt University

Vanderbilt University

  • Nashville, TN

Academic Highlights: Four of Vandy’s ten schools cater to undergrads: the College of Arts and Sciences, the Blair School of Music, the Peabody College of Education and Human Development, and the School of Engineering. In the 2022-23 school year, 87% of course sections contained 19 or fewer students. Of the 70 undergraduate majors, economics, politics and government, and neuroscience are the most popular. The School of Engineering has a strong national reputation as do offerings in biology, economics, education, and music.

Professional Outcomes: Six months after graduating, 96% of the Class of 2021 were employed or in graduate school. The most commonly entered industry was finance followed by technology, consulting, education, and engineering. Alumni can be found in droves at Capital One, Goldman Sachs, Bain & Company, JP Morgan Chase, Citi, and Meta. Among 2022 alumni who directly pursued advanced degrees, the majority enrolled at Vanderbilt followed by Columbia, Harvard, Penn, NYU, and Northwestern.

  • Enrollment: 7,151 (undergraduate); 6,559 (graduate)
  • Cost of Attendance: $89,590

University of Pittsburgh

University of Pittsburgh

  • Pittsburgh, PA

Academic Highlights: Pitt admits freshmen to the Dietrich School of Arts & Sciences, the College of Business Administration, the Swanson School of Engineering, and the School of Nursing. Pitt’s engineering and business schools are top-rated and among the most commonly chosen fields of study. Premed offerings are also top-notch, with majors in the health professions (12%), biology (11%), psychology (9%), and computer science (9%) rounding out the list of most popular majors. Pitt has a strong 13:1 student-to-faculty ratio; 42% of sections have an enrollment of under twenty students.

Professional Outcomes: Within a few months of graduating, 94% of 2022 grads entered full-time employment or full-time graduate or professional school. Engineering, nursing, business, and information sciences majors had 73-86% employment rates while other majors tended to flock to graduate school in large numbers. Employers scooping up the highest number of grads in one recent year included the University of Pittsburgh Medical Center (170), PNC (57), BNY Mellon (36), and Deloitte (19). Median starting salaries fluctuated between $37k-65k depending on major.

  • Enrollment: 20,220 (undergraduate); 9,268 (graduate)
  • Cost of Attendance: $38,034-$43,254 (in-state); $56,400-$66,840 (out-of-state)
  • Median SAT: 1360
  • Median ACT: 31
  • Acceptance Rate: 50%

Reed College

Reed College

  • Portland, OR

Academic Highlights: Twenty-six academic departments collectively offer 38 majors, 13 of which are interdisciplinary. English, math, and physics receive particularly strong praise. The educational experience is highly personalized, and to ensure that the emphasis in the classroom remains on learning for learning’s sake, grades are issued but not shared with students until graduation. The areas in which the highest number of degrees are conferred are the social sciences (22%), biology (11%), the health professions (9%), and the physical sciences (7%).

Professional Outcomes: An examination of Reed’s alumni database reveals that the three most common occupational pathways are business (28%), education (25%), and self-employment (19%). Included among the most frequent current employers are Microsoft Corporation, Kaiser Permanente, Portland Public Schools, Intel Corporation, the US Department of State, National Institutes of Health, and Apple. An incredible amount of Reedies continue their education; Reed was the No. 6 producer of future PhD holders across all academic disciplines.

  • Enrollment: 1,523 (undergraduate); 11 (graduate)
  • Cost of Attendance: $86,040
  • Median SAT: 1410
  • Acceptance Rate: 31%
  • Retention Rate: 86%
  • Graduation Rate: 80%

University of Rochester

University of Rochester

  • Rochester, NY

Academic Highlights: U of R’s 160 undergraduate programs offer a rigorous yet flexible educational experience at a small liberal arts college within a renowned research institution. 66% of sections enroll fewer than 20 students, and an impressive 77% of students are involved in undergraduate research. The Eastman School of Music is one of the best music conservatories in the United States, and the Hajim School of Engineering & Applied Sciences is a top 50 institution. Other strong majors include mathematics, economics, and political science.

Professional Outcomes: Six months after receiving their diplomas, 97% of Class of 2022 grads had achieved positive outcomes with 57% employed and a notably high 38% already pursuing an advanced degree. Popular industries included internet and software (14%), healthcare (14%), and investment banking (6%). Top employers of 2022 grads included Google, Goldman Sachs, Epic Systems, and Deloitte. The average starting salary for the most recent cohort of grads was an impressive $82,325; that figure was over $105k for graduates of the Hajim School of Engineering.

  • Enrollment: 6,767 (undergraduate); 5,430 (graduate)
  • Cost of Attendance: $85,858
  • Acceptance Rate: 39%

Brandeis University

Brandeis University

  • Waltham, MA

Academic Highlights: Brandeis offers 43 majors, the most popular of which are in the social sciences (18%), biology (17%), business (10%), psychology (8%), public administration (8%), and computer science (7%). The student-faculty ratio is 11:1, and 60% of courses contain nineteen or fewer students. Departments with a particularly strong national reputation include economics, international studies, and sociology as well as all of the traditional premed pathways including biology, and chemistry.

Professional Outcomes: Within six months of graduation, 98% of the Class of 2022 had found their way to employment (59%), graduate school (35%), or another full-time activity like travel or volunteer work (4%). Members of the Class of 2022 were hired by Red Hat, Deloitte, Nasdaq, NPR, and McKinsey & Company. The average starting salary for recent grads is $61k. A large contingent of grads elects to continue at Brandeis for graduate school. Many others go to BU, Columbia, Duke, Harvard, and Yale.

  • Enrollment: 3,687
  • Cost of Attendance: $86,242
  • Retention Rate: 91%

Georgetown University

Georgetown University

  • Washington, D.C.

Academic Highlights: The student-faculty ratio is 11:1, and 60% of classes enroll fewer than 20 students. While some classes are a bit larger, only 7% cross the 50-student threshold. Those desiring to join the world of politics or diplomacy are in the right place. The Government and International Affairs programs are among the best in the country. The greatest number of degrees are conferred in the social sciences (38%) followed by business (20%), interdisciplinary studies (8%), and biology (7%).

Professional Outcomes: Within six months of graduating, 75% of members of the Class of 2022 entered the workforce, 19% went directly into a graduate or professional program of study, and 3% were still seeking employment. The Class of 2022 sent massive numbers of graduates to a number of major corporations including JPMorgan Chase (22), Citi (21), BOA (18), Morgan Stanley (16), and EY (10). Those attending grad school stay at Georgetown or flock to other elite schools like Columbia and Harvard.

  • Enrollment: 7,900
  • Cost of Attendance: $85,000
  • Acceptance Rate: 12%

College of William & Mary

College of William & Mary

  • Williamsburg, VA

Academic Highlights: Forty undergraduate programs are available and William & Mary has a 13:1 student-to-faculty ratio. Class sizes are rarely tiny seminars, but 44% do enroll fewer than twenty students, and only 9% contain more than fifty. Among the college’s most notable academic programs are (1) government and (2) international relations, both of which serve as pipelines to Washington, DC, employers. The Mason School of Business is highly regarded in the corporate world. The social sciences (20%) and biology (11%) are also popular.

Professional Outcomes: 52% of the most cohort joined the workforce, and 36% entered graduate school within six months of graduation. Over 500 employers snatched up at least one member of the Tribe. Companies hiring at least four 2020 grads included Accenture, Booz Allen Hamilton, KPMG, and Deloitte. Many of those opting for immediate entry into graduate school stayed at their alma mater and the next most frequented universities included Columbia, Duke, Harvard, Northwestern, and the University of Chicago.

  • Enrollment: 6,797
  • Cost of Attendance: $39,595 (In-State); $63,967 (Out-of-State)
  • Acceptance Rate: 33%

University of Southern California

University of Southern California

Academic Highlights : There are 140 undergraduate majors and minors within the Dornsife College of Arts & Sciences alone, the university’s oldest and largest school. The Marshall School of Business, Viterbi School of Engineering, and programs in communication, the cinematic arts, and the performing arts are highly acclaimed. Popular areas of study are business (22%), social sciences (11%), visual and performing arts (11%), communications/journalism (9%), and engineering (8%). Most courses enroll 10-19 students, and USC does an excellent job facilitating undergraduate research opportunities.

Professional Outcomes: 96% of undergrads experience positive postgraduation outcomes within six months of earning their degree. The top five industries entered were finance, consulting, advertising, software development, and engineering; the median salary across all majors is an astounding $79k. Presently, between 300 and 1,500 alumni are employed at each of Google, Amazon, Apple, Microsoft, KPMG, Goldman Sachs, and Meta. Graduate/professional schools enrolling the greatest number of 2022 USC grads include NYU, Georgetown, Harvard, Stanford, Pepperdine, and UCLA.

  • Enrollment: 20,699 (undergraduate); 28,246 (graduate)
  • Cost of Attendance: $90,921

Case Western Reserve University

Case Western Reserve University

  • Cleveland, OH

Academic Highlights: Sporting a 9:1 faculty-to-student ratio, the university does a nice job keeping classes on the small side, with 50% of course sections capped at nineteen and only 13% of courses having fifty students or more. The Weatherhead School of Management and the Case School of Engineering have stellar reputations within the worlds of employment and academia. Engineering is the most commonly conferred undergraduate degree at 27% followed by biology (15%), computer science (10%), and health professions (8%).

Professional Outcomes: Fifty-six percent of 2022 CWRU graduates head into the world of professional employment upon receiving their diplomas; only 3% were still looking for work six months after leaving the university. More than half of the Class of 2022 enjoyed a starting salary of at least $65,000. The employers of the greatest number of graduates included Accenture, Microsoft, Deloitte, and Google. In 2022, four or more grads were accepted to continue their studies at the likes of NYU, Columbia University, Johns Hopkins University, WashU, and the University of Michigan.

  • Enrollment: 6,017
  • Cost of Attendance: $88,516
  • Median SAT: 1480
  • Acceptance Rate: 27%

University of California, Irvine

University of California, Irvine

Academic Highlights: UCI offers eighty undergrad programs as well as many opportunities for personal connection; 56% of all sections enroll 19 or fewer students and over 60% of students conduct a research project. The most commonly conferred degrees are the social sciences (16%), business (12%), psychology (11%), and biology (9%). The Samueli School of Engineering has a solid reputation as does the Bren School, the only independent computer science school in the UC system. Programs in public health and biological sciences earn very high marks.

Professional Outcomes: Accounting, aerospace, internet and software, K-12 education, real estate, and retail are among the industries attracting the greatest number of Anteaters. Companies employing large numbers of recent grads include Boeing, the Walt Disney Company, Google, EY, and Microsoft. Hundreds of alumni are also found at Kaiser Permanente, Meta, Apple, Edwards Lifesciences, and Deloitte. The median salary is $69,000, with CS grads earning close to $120k right off the bat. UCI has a very strong reputation for premed.

  • Enrollment: 28,661 (undergraduate); 7,275 (graduate)
  • Cost of Attendance: $40,202 (in-state); $72,776 (out-of-state)
  • Acceptance Rate: 26%

We hope you have found our list of the Best Colleges for Biology to be useful and informative as you continue your college search process. We also invite you to check out some of our other resources and tools including:

  • AP Score Calculators 
  • SAT Score Calculator 
  • ACT Score Calculator
  • Best Summer Programs 
  • College List Building Tool
  • Best Colleges by Major

' src=

Andrew Belasco

A licensed counselor and published researcher, Andrew's experience in the field of college admissions and transition spans two decades. He has previously served as a high school counselor, consultant and author for Kaplan Test Prep, and advisor to U.S. Congress, reporting on issues related to college admissions and financial aid.

  • 2-Year Colleges
  • Application Strategies
  • Best Colleges by State
  • Big Picture
  • Career & Personality Assessment
  • College Essay
  • College Search/Knowledge
  • College Success
  • Costs & Financial Aid
  • Data Visualizations
  • Dental School Admissions
  • Extracurricular Activities
  • Graduate School Admissions
  • High School Success
  • High Schools
  • Law School Admissions
  • Medical School Admissions
  • Navigating the Admissions Process
  • Online Learning
  • Private High School Spotlight
  • Summer Program Spotlight
  • Summer Programs
  • Test Prep Provider Spotlight

College Transitions Sidebar Block Image

“Innovative and invaluable…use this book as your college lifeline.”

— Lynn O'Shaughnessy

Nationally Recognized College Expert

College Planning in Your Inbox

Join our information-packed monthly newsletter.

I am a... Student Student Parent Counselor Educator Other First Name Last Name Email Address Zip Code Area of Interest Business Computer Science Engineering Fine/Performing Arts Humanities Mathematics STEM Pre-Med Psychology Social Studies/Sciences Submit

Gravatar Icon

Ecology and Evolutionary Biology Graduate Programs in America

1-25 of 90 results

Yale Graduate School of Arts and Sciences

New Haven, CT •

Yale University •

Graduate School

  • • Rating 4.5 out of 5   2 reviews

Master's Student: The resources at Yale are outstanding. While some of the required courses are slow-moving and less informative, I do have more academic freedom in my second year to the program to take classes that I enjoy across all departments (including data science, statistics, computer science, and law). ... Read 2 reviews

Yale University ,

Graduate School ,

NEW HAVEN, CT ,

2 Niche users give it an average review of 4.5 stars.

Featured Review: Master's Student says The resources at Yale are outstanding. While some of the required courses are slow-moving and less informative, I do have more academic freedom in my second year to the program to take classes that I... .

Read 2 reviews.

Harvard Graduate School of Arts and Sciences

Cambridge, MA •

Harvard University •

  • • Rating 4.56 out of 5   9 reviews

Other: I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student' in the Harvard Graduate School of Arts and Sciences. Through this School, I will be focusing my time at the John A. Paulson school of Engineering & Applied Sciences. Looking forward to wrapping up my final year on campus! ... Read 9 reviews

Harvard University ,

CAMBRIDGE, MA ,

9 Niche users give it an average review of 4.6 stars.

Featured Review: Other says I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student'... .

Read 9 reviews.

Princeton University

Princeton, NJ •

  • • Rating 4.33 out of 5   3 reviews

Master's Student: The best part of the Princeton University mechanical engineering graduate degree is the excellent faculty that teach the courses. They are incredibly knowledgeable and also very willing to help students in office hours or in sponsorship of projects. The worst part of the Princeton University mechanical engineering graduate degree is the lack of structure for the graduate research program which can leave you feeling unsure on the direction of your research. ... Read 3 reviews

PRINCETON, NJ ,

3 Niche users give it an average review of 4.3 stars.

Featured Review: Master's Student says The best part of the Princeton University mechanical engineering graduate degree is the excellent faculty that teach the courses. They are incredibly knowledgeable and also very willing to help... .

Read 3 reviews.

College of Liberal Arts and Sciences

Lesley University •

Graduate School •

CAMBRIDGE, MA

  • • Rating 3 out of 5   2

Florida Institute of Technology

MELBOURNE, FL

  • • Rating 4.61 out of 5   66

Lehigh University

BETHLEHEM, PA

  • • Rating 4.42 out of 5   19

Nicholas School of the Environment

Durham, NC •

Duke University •

  • • Rating 4.86 out of 5   7 reviews

Master's Student: Thus far, my academic experience has been great. I have had access to academic advising sessions and have been able to successfully enroll in my desired classes. An area of improvement would be the required diagnostic exams in calculus and statistics, which cannot be waived even with the successful completion of the undergraduate coursework. ... Read 7 reviews

Duke University ,

DURHAM, NC ,

7 Niche users give it an average review of 4.9 stars.

Featured Review: Master's Student says Thus far, my academic experience has been great. I have had access to academic advising sessions and have been able to successfully enroll in my desired classes. An area of improvement would be the... .

Read 7 reviews.

Alpert Medical School

Providence, RI •

Brown University •

  • • Rating 4 out of 5   1 review

Graduate Student: I have had a great time here. It's really about the people. My fellow classmates are wonderful, brilliant, accomplished people and I am lucky to be here. The faculty are similarly great as are most of the physicians. I have very few complaints but here are 2 fairly big ones: 1. Research isn't emphasized enough. I was told many times first year not to worry about it "yet" but then was never told when I should worry about it. Luckily I found my own research projects but for competitive specialties, you can't act like research isn't a big deal. 2. Clinical grading is so subjective. I've gotten the exact same scores and received honors in one rotation but not another. I've gotten amazing comments but 4/5 scores or good comments but 5/5 scores. Also, having only Honors/Pass/Fail hurts people just out of the Honors range. No one fails. Yet those who barely try "earn" a Pass while those who perform well and try hard could get screwed by one tough evaluator and receive the same Pass. ... Read 1 review

Brown University ,

PROVIDENCE, RI ,

1 Niche users give it an average review of 4 stars.

Featured Review: Graduate Student says I have had a great time here. It's really about the people. My fellow classmates are wonderful, brilliant, accomplished people and I am lucky to be here. The faculty are similarly great as are most... I have very few complaints but here are 2 fairly big ones: 1. Research isn't emphasized enough. I was told many times first year not to worry about it "yet" but then was never told when I should worry about it. Luckily I found my own research projects but... 2. Clinical grading is so subjective. I've gotten the exact same scores and received honors in one rotation but not another. I've gotten amazing comments but 4/5 scores or good comments but 5/5... .

Read 1 reviews.

School of Arts & Sciences - University of Pennsylvania

Philadelphia, PA •

University of Pennsylvania •

University of Pennsylvania ,

PHILADELPHIA, PA ,

  • Find college scholarships

Wiess School of Natural Sciences

Houston, TX •

Rice University •

Blue checkmark.

Rice University ,

HOUSTON, TX ,

Washington University in St. Louis - Arts & Sciences

St. Louis, MO •

Washington University in St. Louis •

Washington University in St. Louis ,

ST. LOUIS, MO ,

Guarini School of Graduate and Advanced Studies

Hanover, NH •

Dartmouth College •

  • • Rating 5 out of 5   1 review

Alum: I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary learning. Guarini is well integrated within the College, which provides a unique world-class learning experience. Guarini went beyond my expectations and made this experience the best I had and will ever have in my life. The only thing I would like changed is having a GRAD diploma in Latin instead of English to keep up with College tradition. ... Read 1 review

Dartmouth College ,

HANOVER, NH ,

1 Niche users give it an average review of 5 stars.

Featured Review: Alum says I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary... .

Biological Sciences Division - University of Chicago

Chicago, IL •

University of Chicago •

Master's Student: So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. ... Read 1 review

University of Chicago ,

CHICAGO, IL ,

Featured Review: Master's Student says So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. .

Marshall School of Business

Los Angeles, CA •

University of Southern California •

  • • Rating 4.71 out of 5   17 reviews

Current Master's student: Marshall is a fantastic business program. So far, my academic experience has been nothing short of fantastic. I have never enjoyed school before, but almost every class I have learned new, applicable, and incredibly helpful techniques that I apply in my job every day. Most of the professors are very helpful, and passionate about their subjects. Marshall has clubs for almost every industry, so depending on your industry of interest, Marshall has clubs to prepare you for interviews and get you great exposure, and they're a great way to network as well. If your industry is not represented, you have the opportunity to create one. Marshall is still relatively underrepresented in minority categories like women and people of color, but from my understanding, they are working to improve those numbers. ... Read 17 reviews

University of Southern California ,

LOS ANGELES, CA ,

17 Niche users give it an average review of 4.7 stars.

Featured Review: Current Master's student says Marshall is a fantastic business program. So far, my academic experience has been nothing short of fantastic. I have never enjoyed school before, but almost every class I have learned new,... Marshall has clubs for almost every industry, so depending on your industry of interest, Marshall has clubs to prepare you for interviews and get you great exposure, and they're a great way to... Marshall is still relatively underrepresented in minority categories like women and people of color, but from my understanding, they are working to improve those numbers. .

Read 17 reviews.

Cornell University College of Agriculture and Life Sciences

Ithaca, NY •

Cornell University •

Cornell University ,

ITHACA, NY ,

  • Sponsored Find Student Loan Options
  • Genetics and Genomics Graduate Programs
  • Physiology and Pathology Graduate Programs

Rackham School of Graduate Studies

Ann Arbor, MI •

University of Michigan - Ann Arbor •

  • • Rating 4.8 out of 5   5 reviews

Master's Student: The Landscape Architecture program at UMich School for Environment and Sustainability is rooted in advancing sustainable design and ecological function, rather than pure aesthetics. We have some amazing faculty very dedicated to this mission, some of whom are legends within the field. This program attracts and retains students who are diverse, passionate, friendly and helpful, and the experience at this school has been very rewarding. The curriculum is challenging but thought provoking, and everyone in the studios is happy and willing to help, fostering a warm sense of comradery and support. ... Read 5 reviews

University of Michigan - Ann Arbor ,

ANN ARBOR, MI ,

5 Niche users give it an average review of 4.8 stars.

Featured Review: Master's Student says The Landscape Architecture program at UMich School for Environment and Sustainability is rooted in advancing sustainable design and ecological function, rather than pure aesthetics. We have some... .

Read 5 reviews.

Tufts University School of Medicine

Boston, MA •

Tufts University •

  • • Rating 4.84 out of 5   25 reviews

Doctoral Student: I am just starting but have looked through some modules. It seems like it is going to be a great experience and I have visited the campuses a couple of times. ... Read 25 reviews

Tufts University ,

BOSTON, MA ,

25 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says I am just starting but have looked through some modules. It seems like it is going to be a great experience and I have visited the campuses a couple of times. .

Read 25 reviews.

College of Arts & Sciences - University of North Carolina at Chapel Hill

Chapel Hill, NC •

University of North Carolina at Chapel Hill •

University of North Carolina at Chapel Hill ,

CHAPEL HILL, NC ,

College of Natural Sciences

Austin, TX •

University of Texas - Austin •

University of Texas - Austin ,

AUSTIN, TX ,

College of Arts and Sciences - Lehigh University

Bethlehem, PA •

Lehigh University •

Lehigh University ,

BETHLEHEM, PA ,

University of Florida College of Agricultural and Life Sciences

Gainesville, FL •

University of Florida •

  • • Rating 4.67 out of 5   6 reviews

Alum: This is a program intended for people who have a purpose. Many enrolled already work full time and want a higher degree to boost payscale or in the case of teachers, allow them to teach higher education programs or develop new ones. For individuals seeking further education, this can be a great stepping stone, however be advised you need to work and seek your path, it will not be handed to you and there is little guidance without effort. Much of the course work is as described- intended for medical microbiology. Some classes are highly outdated and most geared towards academia not industry. I liked the ability to customize some of the coursework but was disappointed a dreadful career seminar I did not want to take was later removed as a requirement without compensation. I would have happily taken a credit even as an audit to learn. Be advised it can be challenging to develop relationships with professors and standout amongst large numbers of students online. ... Read 6 reviews

University of Florida ,

GAINESVILLE, FL ,

6 Niche users give it an average review of 4.7 stars.

Featured Review: Alum says This is a program intended for people who have a purpose. Many enrolled already work full time and want a higher degree to boost payscale or in the case of teachers, allow them to teach higher... .

Read 6 reviews.

College of Liberal Arts & Sciences - University of Illinois

Urbana, IL •

University of Illinois Urbana-Champaign •

University of Illinois Urbana-Champaign ,

URBANA, IL ,

Odom School of Ecology

Athens, GA •

University of Georgia •

University of Georgia ,

ATHENS, GA ,

College of Letters and Science - UC Santa Barbara

Santa Barbara, CA •

University of California - Santa Barbara •

University of California - Santa Barbara ,

SANTA BARBARA, CA ,

BYU College of Physical and Mathematical Sciences

Provo, UT •

Brigham Young University •

Brigham Young University ,

PROVO, UT ,

College of Food, Agricultural and Natural Resource Sciences - University of Minnesota Twin Cities

Saint Paul, MN •

University of Minnesota Twin Cities •

University of Minnesota Twin Cities ,

SAINT PAUL, MN ,

College of Biological Sciences - University of Minnesota Twin Cities

University of washington college of arts & sciences.

Seattle, WA •

University of Washington •

University of Washington ,

SEATTLE, WA ,

Lesley University

  • • Rating 4.52 out of 5   31

College of Arts and Sciences - University of Wyoming

University of Wyoming •

LARAMIE, WY

  • • Rating 5 out of 5   4

Showing results 1 through 25 of 90

Department of Biology

From Molecules to Ecosystems

Portland State University's Biology Department includes over 21 faculty, 1,200 undergraduate majors, and over 65 graduate students and post-docs. We take an integrative approach to biology, encompassing all levels of biological organization from molecules to ecosystems.

Our program prepares students for careers in biological research, teaching, health sciences, biotechnology, conservation biology and wildlife management, forestry, and other applied fields.

Human cell

Grad Applications Due Feb. 1

Our multi-disciplinary faculty offer admitted graduate students the opportunity to conduct cutting edge research, gain professional teaching experience, and pursue an academically rigorous program which fosters professional research development.

Our Programs and Events

Students in science lab with instructor up front

Undergraduate Studies

two people looking at a computer

Graduate Studies

Scientists working in Antarctica

Weekly Seminars

Biology news.

paper

Currently Accolades: Published/Exhibited for May 6, 2024

Portland State University Day of Giving

Help Our Students Thrive: Join CLAS for PSU’s Spring Day of Giving on April 17

Graphic icon of three people

CLAS Grad Student Accolades: Presentations for 2023

Best Global Universities for Biology and Biochemistry

The best universities for biology and biochemistry show strength in producing research in the field and have top-notch reputations. This field includes many subjects, such as anatomy, physiology and pathology, which all share a core mission of analyzing the science of life. These are the world's top universities for biology and biochemistry. Read the methodology »

To unlock more data and access tools to help you get into your dream school, sign up for the  U.S. News College Compass !

Here are the best global universities for biology and biochemistry

Harvard university, massachusetts institute of technology (mit), stanford university, university of california berkeley, university of california san francisco, university of cambridge, university of california san diego, university of washington seattle, university of oxford, johns hopkins university.

See the full rankings

  • Clear Filters

best phd programs for biology

  • # 1 in Best Universities for Biology and Biochemistry
  • # 1 in Best Global Universities

Founded in 1636, Harvard University is the oldest higher education institution in the U.S. The bulk of Harvard's... Read More

best phd programs for biology

  • # 2 in Best Universities for Biology and Biochemistry
  • # 2 in Best Global Universities

Massachusetts Institute of Technology, founded in 1861, is located in Cambridge, Massachusetts, near Boston. Around... Read More

best phd programs for biology

  • # 3 in Best Universities for Biology and Biochemistry
  • # 3 in Best Global Universities

Stanford University was founded in 1885 and is located in California’s Bay Area, around 30 miles south of San Francisco... Read More

best phd programs for biology

  • # 4 in Best Universities for Biology and Biochemistry
  • # 4 in Best Global Universities

The University of California—Berkeley is situated roughly 15 miles from San Francisco in what is known as the Bay Area... Read More

best phd programs for biology

  • # 5 in Best Universities for Biology and Biochemistry
  • # 16 in Best Global Universities  (tie)

The University of California—San Francisco is a public institution that was founded in 1864. The health sciences-focused... Read More

best phd programs for biology

  • # 6 in Best Universities for Biology and Biochemistry
  • # 8 in Best Global Universities

The University of Cambridge, located around 60 miles north of London, traces its history back to 1209. Around 19,000... Read More

best phd programs for biology

  • # 7 in Best Universities for Biology and Biochemistry
  • # 20 in Best Global Universities

The University of California—San Diego is a public institution that was established in 1960. It is located in the La... Read More

best phd programs for biology

  • # 8 in Best Universities for Biology and Biochemistry
  • # 6 in Best Global Universities

The University of Washington is a public institution that was founded in 1861. The school's oldest and largest campus in... Read More

best phd programs for biology

  • # 9 in Best Universities for Biology and Biochemistry
  • # 5 in Best Global Universities

The exact date of the University of Oxford’s founding is unknown, but the school traces its roots back to at least 1096... Read More

best phd programs for biology

  • # 10 in Best Universities for Biology and Biochemistry
  • # 10 in Best Global Universities

Johns Hopkins University is a private institution that was founded in 1876. The school has campuses located in and... Read More

IMAGES

  1. 2023 Best Online PhD in Biology Programs [Doctorate Guide]

    best phd programs for biology

  2. The 10 Best Biology PhD Programs

    best phd programs for biology

  3. 2023 Best Online PhD in Biology Programs [Doctorate Guide]

    best phd programs for biology

  4. 2024 Best Online PhD in Biology Programs [Doctorate Guide]

    best phd programs for biology

  5. 2023 Best Online PhD in Biology Programs [Doctorate Guide]

    best phd programs for biology

  6. Best Molecular Biology PHD Programs

    best phd programs for biology

VIDEO

  1. Pursue a PhD at the frontiers of biomedical sciences

  2. Fully Funded PhD Scholarship at the Institute of Science and Technology Austria (ISTA)

  3. How to avoid Toxic PhD Labs: Strategies for Research Success #phd #research #drjyotibala

  4. Phd admission 2024 !how to take admission and interview preparation and apply for best institutes

  5. PhD Student’s Story: From Pakistan to America

  6. Youngest PhD Holder in Bangladesh

COMMENTS

  1. Ph.D. Program

    The training for a Ph.D. in Biology is focused on helping students achieve their goals of being a successful research scientist and teacher, at the highest level. Students work closely with an established advisor and meet regularly with a committee of faculty members to facilitate their progress. The Biology Ph.D. program is part of the larger ...

  2. 2023-2024 Top Biology Graduate Programs

    My intended Advanced Academic Program is the accelerated (2 semester), dual-modality, 40-credit M.S. in Biotechnology, Biodefense concentration. All of the anticipated course subjects are diverse and there's even a customizable core lab course on campus (at least until Summer 2024).

  3. Best Biological Sciences Programs

    University of California--San Francisco. San Francisco, CA. #9 in Biological Sciences (tie) Save. 4.7. A graduate degree in biological sciences gives students a broad base of knowledge in the life ...

  4. Best Biophysics / Structural Biology Programs

    Ranked in 2022, part of Best Science Schools. Biochemistry, biophysics and structural biology courses combine principles of biology, chemistry and physics and are often heavily research-based.

  5. Best Molecular Biology Programs

    Best Molecular Biology Programs. Ranked in 2022, part of Best Science Schools. Molecular biologists are responsible for research in many disciplines, including the study of disease prevention, DNA ...

  6. Molecular and Cellular Biology

    The Molecules, Cells, and Organisms PhD program in the Department of Molecular and Cellular Biology provides a view of the broad range of the constantly evolving world of scientific experience. In this interdisciplinary program, you will interact with students and faculty who have diverse backgrounds in chemistry, marine biology, computational ...

  7. Systems, Synthetic, and Quantitative Biology

    You can find degree program-specific admissions requirements below and access additional guidance on applying from the Systems, Synthetic, and Quantitative Biology PhD Program. Academic Background Applicants typically have a background in biology, physics, chemistry, computer science, engineering, or mathematics and work to forge a new approach ...

  8. Doctoral Program

    The Department of Molecular & Cell Biology at the University of California, Berkeley offers a Ph.D. program focused on the molecular mechanisms inherent to life. This program integrates research with a modern training curricula, teaching, and career mentorship. Our Department is highly interdisciplinary - comprising the Divisions of Cell Biology, Development & Physiology, Immunology and ...

  9. Biology, PhD < University of Pennsylvania

    Biology, PhD. The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student's thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology ...

  10. Top Biology Doctorate Degrees & Graduate Programs 2024+

    The PhD in Biology degree is an in-depth, research degree and the highest level of education in the field. In most biology PhD programs, students choose a specific area in the biological sciences to anchor their studies. Some of the options might include Cell and Molecular Biology, Ecology and Evolution, Neurobiology and Computational Biology.

  11. PhD in Molecular Microbiology and Immunology

    MMI PhD students hone their scholarship, research and professional skills. Course and laboratory work can address problems in microbial pathogenesis, immunology, disease transmission, and diseases related to malaria, mosquito and arboviral biology. Students can choose to complete the traditional MMI PhD program or the MMI PhD program ...

  12. Systems, Synthetic, and Quantitative Biology PhD Program

    The Systems, Synthetic, and Quantitative Biology PhD Program aims to explain how higher level properties of complex biological systems arise from the interactions among their parts. This field requires a fusion of concepts from many disciplines, including biology, computer science, applied mathematics, physics and engineering. ...

  13. Doctor of Philosophy (PhD) in Biology

    The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of ...

  14. Best Cell Biology Programs

    Ranked in 2022, part of Best Science Schools. With a graduate degree in cell biology, scientists may find jobs in research institutions, colleges and universities, pharmaceutical companies and ...

  15. Best 308 Biology PhD Programmes in United States 2024

    308 Biology PhDs in United States. Molecular and Cell Biology. University of Maryland Baltimore County (UMBC) Baltimore Highlands, Maryland, United States. Biological Sciences - Cell Biology and Molecular Genetics. Kent State University. Kent, Ohio, United States. Biological and Agricultural Engineering.

  16. PhD in Biochemistry and Molecular Biology

    The Biochemistry and Molecular Biology PhD program prepares students for a range of biomedical and health sciences careers, including in academia, industry, policy, and beyond. Visit the Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries. Sample Careers. Research Scientist ...

  17. The 10 Best Biology PhD Programs

    9. University of Wisconsin-Madison. Madison, WI. 10. University of Michigan, Ann Arbor. Ann Arbor, MI. Show More. This is a ranking of the 10 Best Biology PhD Programs in the US. A PhD can potentially prepare students with a passion for biology for many different careers.

  18. Graduate Studies

    The graduate program in Developmental Biology leads to the PhD degree. We offer a joint training program in Genetics and Developmental Biology. The department also participates in the Medical Scientists Training Program (MSTP) in which individuals are candidates for both the MD and PhD degrees. Students are required to complete at least six ...

  19. 50 Best Colleges for Biology

    Best Colleges for Biology - we reveal the 50 best biology colleges in the United States in 2024. Profiles of each school are included. ... 48% were enrolled in master's programs, 23% in PhD programs, 13% in law school, and 8% in med school. The top twenty graduate schools attended, by volume, in the last five years make an exclusive list ...

  20. Top Biology and Biochemistry Universities in United States

    Italy. Japan. Netherlands. See the US News rankings for Biology and Biochemistry among the top universities in United States. Compare the academic programs at the world's best universities.

  21. 2023-2024 Top Ecology and Evolutionary Biology Graduate Programs

    About This List. Explore ecology and evolutionary biology graduate programs and graduate schools offering ecology and evolutionary biology degrees. Compare graduate ecology and evolutionary biology programs with government statistics and graduate student reviews. Find the best ecology and evolutionary biology graduate schools for you.

  22. Biology

    Portland State University's Biology Department includes over 21 faculty, 1,200 undergraduate majors, and over 65 graduate students and post-docs. We take an integrative approach to biology, encompassing all levels of biological organization from molecules to ecosystems. Our program prepares students for careers in biological research, teaching ...

  23. 2024 Best Graduate Science Schools

    Cell Biology. Ecology / Evolutionary Biology. Genetics / Genomics / Bioinformatics. Immunology / Infectious Disease. Microbiology. Molecular Biology. Neuroscience / Neurobiology. See the best ...

  24. Top Biology and Biochemistry Schools in the World

    Germany. India. Italy. Japan. Netherlands. See the US News rankings for the world's top universities in Biology and Biochemistry. Compare the academic programs at the world's best universities.